Herbs annual, biennial, or short-lived perennial. Trichomes short-stalked, 1-3-forked, mixed with simple ones. Stems procumbent, decumbent, or ascending, rarely erect, few to numerous from base. Basal leaves petiolate, rosulate only early, later deciduous, 1-or 2-pinnatisect. Cauline leaves petiolate or uppermost subsessile, similar to basal ones. Racemes bracteate throughout or only basally, elongated in fruit. Fruiting pedicels suberect or divaricate. Sepals oblong, suberect, base of lateral pair not saccate. Petals white, longer than sepals; blade obovate, apex rounded; claw subequaling sepals. Stamens 6, slightly tetradynamous; filaments not dilated at base; anthers ovate, obtuse at apex. Nectar glands 4, lateral, 1 on each side of lateral stamen; median glands absent. Ovules 18-46 per ovary. Fruit dehiscent silicles or rarely siliques, broadly to narrowly oblong, obovate, or orbicular, strongly angustiseptate, sessile; valves papery, with a prominent midvein, glabrous or pubescent, strongly keeled, slightly torulose, flat or twisted; replum rounded; septum complete or rarely perforated; style distinct, often less than to 1 mm; stigma capitate, entire. Seeds uniseriate, wingless, oblong, plump; seed coat coarsely reticulate, not mucilaginous when wetted; cotyledons incumbent.