Annuals or perennials, 30–200 cm (taprooted). Stems decumbent to erect, glabrous, hirsute, or puberulent. Leaves mostly cauline; mostly alternate; petioles 1–7 cm; blades deltate-ovate, lance-ovate, or ovate, 2.5–14 × 1.8–13 cm, bases cordate to truncate or broadly cuneate, margins subentire to serrate, abaxial faces glabrate to hispid, not gland-dotted. Heads 1–3. Peduncles 9–50 cm. Involucres hemispheric, 10–22 mm diam. Phyllaries 20–30, lanceolate, 8–17 × 1–3 mm, apices acute to long-attenuate, abaxial faces glabrous or ± hispid, not gland-dotted. Paleae 7.5–8 mm, apices 3-toothed (middle teeth acuminate, usually glabrous or hispid, sometimes ± villous or bearded). Ray florets 11–20; laminae 12–23 mm. Disc florets 30+; corollas 4.5–5 mm, lobes usually reddish, sometimes yellow; anthers dark, appendages dark (style branches usually reddish, rarely yellow). Cypselae 2.5–3.2 mm, glabrous or sparsely hairy; pappi of 2 lanceolate or lance-linear scales 1.2–2.5 mm.