Heywoodia Sim

Genus

Angiosperms > Malpighiales > Phyllanthaceae

Characteristics

Dioecious evergreen tree lacking indumentum. Leaves alternate, petiolate, peltate in the seedling stage and on sucker-shoots, otherwise basally attached, stipulate, simple, entire, symmetrical at the base, coriaceous, penninerved. Flowers fasciculate, the males densely so, and sessile, the females in 1–3-flowered cymes and shortly pedicellate, axillary. Male flowers: bracts 4, free, imbricate, broad, concave, unequal; sepals 3, free, imbricate, broad, concave, unequal, in size and texture intermediate between the bracts and the petals; petals 5, free, imbricate, concave, subequal; disc peripheral, crisped-lobulate, invaginated amongst the filaments of the outer whorl of stamens, fleshy; stamens (8–)10–11(–12), in 2 whorls, filaments free, anthers erect, slightly recurved, dorsifixed, introrse, thecae parallel, longitudinally dehiscent; pistillode minute, trifid. Female flowers: bracts, sepals and petals ± as in the ♂; 1–2 bracteoles also present; disc ± annular, slightly lobulate, shallowly cupular; staminodes 6–8, filiform; ovary 4–5-locular, with 2 ovules per locule; stigmas 4–5, sessile, bilobed. Fruit slightly depressed-subglobose, shallowly 8–10-lobed and 4–5-ridged, tardily dehiscent, leaving a columella; pericarp thinly fleshy, smooth, becoming rugulose when dried; endocarp bony. Seeds solitary per locule by abortion, ecarunculate, obliquely ovoid, rounded-triangular in section, ± smooth; endosperm papery; embryo minute, cotyledons broad, flat.
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Male flowers: sepals 3, free, imbricate, concave, unequal; petals 5, free, imbricate, concave, subequal; disk extrastraminal, lobulate, invaginated amongst the outer stamens, fleshy; stamens (8)10–11(12), in 2 whorls, filaments free, anthers dorsifixed, introrse, thecae parallel, longitudinally dehiscent; pistillode (non-functional ovary) minute, trifid.
Female flowers: bracteoles 1–2, sepals and petals similar to those of the male; disk annular, slightly lobulate, shallowly cupular; staminodes 6–8, filiform; ovary 4–5-locular, with 2 ovules per loculus; stigmas 4–5, sessile, 2-lobed.
Fruit depressed-subglobose, shallowly 8–10-lobed, tardily septicidally dehiscent, leaving a columella; pericarp thinly fleshy, smooth, becoming rugulose on drying; endocarp bony.
Flowers axillary, fasciculate, the males densely so, sessile, bracteate, the females in 1–3-flowered cymes, shortly pedicellate; bracts 4, free, imbricate, concave, unequal.
Leaves alternate, petiolate, sometimes peltate in juvenile foliage, stipulate; blades simple, entire, basally symmetrical, coriaceous, penninerved.
Seeds solitary per loculus by abortion, ecarunculate; endosperm papery; embryo minute; cotyledons broad, flat.
Dioecious, completely glabrous evergreen tree.
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Foliage retention evergreen
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Hardiness (USDA) 9-12

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Cultivation

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Distribution

Heywoodia world distribution map, present in Kenya, Mozambique, eSwatini, Tanzania, United Republic of, and Uganda

Identifiers

LSID urn:lsid:ipni.org:names:15501-1
WFO ID wfo-4000017867
COL ID 8VY8L
BDTFX ID -
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Heywoodia

Lower taxons

Heywoodia lucens