Leaves ovate or elliptic to broadly lanceolate, acuminate, rounded to cuneate and often unequal at base, petiolate.
Ovary subspherical, slightly compressed laterally, unicarpellate; style indistinct or short, stigma capitate.
Sepals 4, one free, 3 united with centre one slightly larger, accrescent at fruiting.
Fruit lenticulate, sometimes slightly asymmetric, pericarp reticulately wrinkled.
Stamens 4–13; filaments filiform; anthers dorsifixed.
Flowers hermaphrodite, more or less zygomorphic.
Inflorescences terminal and axillary racemes.
Herbs, sometimes woody at base.
Seed discoidal; testa black.
Bracts single, caudate.
Bracteoles in pairs.