Plants medium-sized to large, in creeping mats or patches. Stems green, loosely complanate-foliate; hyalodermis and sclerodermis absent, central strand present. Leaves broad, dorsal leaves often broader, more symmetric than lateral leaves; margins plane, entire, unbordered; apex rounded-obtuse to sharply or broadly acute, flat; ecostate; laminal cells large, lax, smooth, uniform across insertion and base or somewhat shorter than medial. Specialized asexual reproduction by filaments among apical leaf rhizoids, filaments unbranched, 1-seriate, subpapillose. Sexual condition autoicous or sometimes dioicous; perigonia on stem, gemmate. Seta red to reddish or blackish, elongate, smooth. Capsule suberect to pendulous, smooth; exothecial cells strongly collenchymatous; annulus distinct; peristome dark red, diplolepidous; exostome unfurrowed, with zigzag median line; endostome basal membrane high, segments extending barely beyond exostome teeth, ± perforate, cilia rudimentary or absent. Calyptra somewhat lobed at base, multistratose at middle, smooth, naked.
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