Stems creeping, 1-3 mm wide across leafy stem, sympodial, remotely and irregularly pinnate to regularly 1-or 2-pinnate; paraphyllia many, base multiseriate, branches 1-or 2-seriate. Stem leaves erect to spreading, heteromallous, often remote, broadly ovate, ovate-deltoid, or ovate, strongly plicate (often obscuring costa), not rugose, 0.8-2.5 mm; base , not to somewhat decurrent; margins serrulate to nearly entire basally, spinose-serrate to serrate distal 2/3; apex acute to acuminate; costa single, double, or rarely triple, sometimes 2-fid, 1/3-3/4 leaf length; alar cells not differentiated; laminal cells smooth. Branch leaves ovate to lanceolate; . Capsule inclined to horizontal; operculum conic; exostome teeth irregularly cross striolate to somewhat reticulate proximally; endostome segments narrowly perforate.
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