Plants in medium-sized to large wefts, often extensive sheets. Stems creeping to ascending or erect, sympodial or monopodial, irregularly branched to regularly pinnate, innovations often ascending-arching; paraphyllia sometimes present, branched, cells elongate; pseudoparaphyllia foliose; rhizoids few, at base of stems and apices of attenuate branches. Stem leaves very broadly ovate to lanceolate; margins serrate to serrulate, sometimes nearly entire; apex acuminate or sometimes acute to rounded; costa usually double, 1/4-2/3 leaf length, sometimes single or nearly ecostate; alar cells sometimes differentiated; medial laminal cells narrowly elliptic to linear-flexuose, prosenchymatous. Branch leaves usually smaller, narrower; margins more sharply toothed; costa stronger. Sexual condition dioicous. Seta elongate, smooth. Capsule usually inclined to horizontal; peristome double; exostome usually reticulate proximally, papillose distally; endostome basal membrane usually high, segments broad, perforate along keel by narrow slits or broadly elliptic gaps, cilia 1-4 (absent in Leptohymenium). Calyptra cucullate, smooth, naked. Spores spheric, finely papillose to nearly smooth.
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