Stems creeping to ascending-arcuate, 2-4 mm wide across leafy stem, sympodial [sometimes monopodial], irregularly branched to regularly 1-or 2-pinnate; paraphyllia many, base 1-or 2-seriate, branches 1-seriate. Stem leaves spreading to squarrose-recurved, heteromallous, not crowded, ovate to broadly ovate, moderately plicate at base (not obscuring costa), sometimes rugose in acumen, (1.1-)2-3 mm; base , not decurrent; margins serrate to serrulate; apex gradually to abruptly acuminate; costa double, weak or to mid leaf; alar cells not differentiated; laminal cells smooth. Branch leaves ovate to elliptic-lanceolate; . Capsule inclined to horizontal; operculum conic, stoutly rostrate; exostome teeth irregularly cross striolate proximally; endostome segments broadly perforate.