Stems creeping to ascending or erect, 2-8 mm wide across leafy stem, monopodial, irregularly and remotely branched to subpinnate; paraphyllia absent. Stem leaves erect-spreading to squarrose, squarrose-recurved, or falcate-secund, crowded or not, broadly ovate, ovate-triangular, or ovate-lanceolate, sometimes plicate proximally, sometimes rugose, 2.1-4.9 mm; base not or slightly decurrent; margins serrulate to nearly entire proximally, serrulate distally; apex acuminate; costa double, short or to 2/3 leaf length; alar cells not differentiated, or shorter, wider than basal cells; laminal cells smooth or coarsely prorate. Branch leaves ovate to narrowly lanceolate. Capsule horizontal; operculum conic; exostome teeth cross striolate to reticulate; endostome segments broadly perforate.