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Female flowers axillary, solitary or paired, or in few-flowered terminal racemes; pedicels slender; sepals 4–6(9), usually free, open or imbricate, narrow, caducous; petals absent; disk absent; ovary 2-locular, compressed perpendicularly to the septum, with 2 apical pendulous anatropous ovules per locule; styles 2, free, elongate, simple with smooth or papillose stigmas.
Fruit stipitate or not, 2-coccous, flattened, winged, septifragally dehiscent or not; pericarp somewhat crustaceous; endocarp thinly membranous; columella persistent.
Seeds usually 1 per coccus, flat, winged or not; testa thin, striate, shiny; endosperm sparse; cotyledons broad, flat, thin; radicle elongate.
Male inflorescences often precocious, axillary, solitary or paucifasciculate, spicate or subpaniculate, densely flowered; bracts minute.
Leaves alternate, shortly petiolate, stipulate, simple, entire, penninerved, acarodomatiiferous, gland-dotted beneath.
Dioecious, deciduous trees or shrubs.
Characters of the family.
Stipules deciduous.
Indumentum simple.
