Plants rupestral or rarely terrestrial, medium-sized. Rhizomes shortly creeping, ascending, or long creeping, stout, densely covered with scales and hairs or only scales; scales ovate-lanceolate or rarely linear-lanceolate. Fronds approximate, subclustered, or remote; stipe stramineous, base swollen into shuttle form and concealed in scales (Hypodematium), or articulate to rhizome, glabrous (Leucostegia); lamina deltoid or ovate-oblong to pentagonal-ovate, 3-pinnate to finely 4-pinnate-pinnatifid, anadromous, herbaceous or papery, both surfaces, rachises, and costules frequently covered with acicular and/or glandular hairs, rarely glabrous. Veins free, pinnate, branches simple or forked. Sori orbicular; indusia large, reniform, ± covered with acicular or glandular hairs, rarely glabrous. Spores ellipsoid, monolete. x = 41.