Leucostegia C.Presl

Rhizome bearing scales and hairs or only scales. Roots borne on all sides of rhizome; the vascular structure is a dorsiventral dictyostele with a thick dorsal and a thick ventral strand, with alternating leaf gaps and bud traces branching from stelar strands at the base of each leaf gap with two simple leaf traces; scales glabrous or bearing multiseptate hairs, basifixed, base not cordate. Stipes articulated at the base to phyllopodia, grooved or not, glabrous. Lamina compound, tri-or quadripinnate (rarely in small plants bipinnate) towards base and in the middle part, deltoid and broadest towards base, glabrous (sometimes minute hairs present), entire to pinnatilobed (often fertile leaves more strongly dissected). Pinnae deltoid or narrowly triangular. Pinnules of at least the larger pinnae anadromous. Pinnules or pinna lobes narrowly ovate. Rhachis adaxially side grooved. Leaf axes glabrous (sometimes a few minute hairs present). Veins in ultimate lobes simple, not reaching the margin. False veins not present. Sori indusiate, frequently single on a segment, terminal on the veins. Indusium scaly, either attached at the narrow, cordate base only, or attached at the base and only part of the sides.