Herbs, erect or spreading, unarmed, hairs simple, often glandular. Leaves ovate or elliptic, entire or mostly toothed, glabrate or viscid, membranaceous, often with large minor leaves. Inflorescence solitary, a slender peduncled umbel situated at a dichotomy of the stem. Flowers with the calyx lobes folded over the bud, becoming erect or outward-curved, deltoid-obtuse, accrescent, spreading, and subtending but not enclosing the fruit; corolla rotate or broadly campanulate, lobed to about halfway, the lobes broad, deltoid; filaments exserted, slender, inserted near the base of the corolla, mostly pubescent, anthers short, dehiscing longitudinally; ovary glabrous, viscid, situated on a disc, 2-loculed, the style slender, the stigma small. Fruit a globose, many-seeded, mucilaginous berry, the surface sometimes viscid; seeds compressed-lenticular, the embryo curved around the endosperm at the periphery of the seed.