Juncaceae Juss.

Rushes (en)

Family

Angiosperms > Poales

Characteristics

Perennial, rhizomatous (usually), stoloniferous or tufted, seldom annual herbs; silica bodies ± absent; glabrous or with hairs on leaf margins, rarely the abaxial leaf surface with simple, many-celled hairs. Culms simple, upright to ascending, rarely prostrate, usually terete (rarely flattened), solid or hollow, leafless or leaf-bearing. Leaves alternate, rarely distichous (not in Australia), 3-ranked; sheath closed or open, often auriculate; blades simple, laterally compressed or terete or reduced to a mucro surmounting the leaf-sheaths (cataphylls, in some texts referred to as ‘basal sheathing bracts’), solid or hollow with internal septa; margins usually entire; stipules absent. Inflorescence terminal or pseudolateral, mainly of cymes contracted into 1–many ± stalked clusters or expanded into panicles, rarely reduced to a single terminal or lateral flower; bract(s) subtending the inflorescence 1 or more; bracteoles (prophylls) subtending solitary flower 0–2(rarely more), translucent, reduced. Flowers bisexual, rarely unisexual and plants dioecious or monoecious, actinomorphic, inconspicuous. Tepals 6, free, in 2 whorls of 3, imbricate, glume-like, persistent. Stamens 3–6, if 6 then in two whorls of 3, filaments free, opposite the tepals; anthers basifixed, tetrasporangiate, dithecal, dehiscing by longitudinal lateral slits, connective rarely with an apical mucro (not in Australia); pollen shed in tetrads. Gynoecium of 3 connate carpels. Ovary superior, placentation axile or parietal, or occasionally basal (Luzula), 1-locular (not in Australia), 3-septate (not in Australia) or 3-locular; style 1 (or rarely 3 and free), apical, simple; stigma 3, terete and of equal diameter throughout or sometimes tapering distally, twining with adaxial papillae. Ovules numerous (rarely 3). Fruit a loculicidal capsule. Seeds 3–many, minute, sometimes with tail-like apical and or basal appendages, often with distinct ornamentation, sometimes mucilaginous; embryo straight, embedded in a starchy endosperm.
More
Herbs, very rarely shrublike, perennial or annual, tufted or with erect or creeping rhizome. Stems erect, terete or laterally flattened. Leaves occasionally reduced to a bladeless or nearly bladeless sheath at base of stem (cataphyll); leaf sheath often shortly prolonged on both sides into a pair of auricles at leaf blade juncture; leaf blade of basal and cauline leaves usually linear or filiform, grasslike and flat, or terete, glabrous except for pilose margin on some grasslike blades. Inflorescence a panicle, corymb, or 1-flowered. Flowers bisexual or unisexual and plants dioecious, mostly wind pollinated, regular, usually small, usually 1-or 2-bracteolate at base. Perianth segments (3 or)6, in (1 or)2 whorls, free, usually greenish to brownish or blackish, rarely white or yellowish, glumelike. Stamens 3 or 6, if 6 then 3 opposite outer perianth segments; filaments thin; anthers basifixed, 2-loculed, dehiscing by longitudinal slits; pollen grains in tetrads. Ovary superior, 1-loculed, or divided by 3 septa and 3-loculed, or incompletely septate; ovules 3 and inserted at base of ovary, or numerous and biseriate on 3 parietal placentas. Stigmas 3, papillose. Fruit a capsule, 1--3-valved, loculicidal. Seeds globose, ovoid, or fusiform, small, sometimes appendaged; appendage caudate; embryo straight, minute, enclosed by fleshy endosperm.
Annual or perennial herbs, rarely shrubs. Roots fibrous. Stems erect, cylindric or rarely compressed, naked or leafy, sometimes rhizomatous or stoloniferous, and then horizontal or ascending. Leaves grass-like or cylindric, sheathing at the base, sometimes reduced to cataphylls; sheaths open and sometimes auriculate, or closed. Inflorescence terminal, often pseudo-lateral, compound or rarely simple with one flower, umbellate, paniculate, the flowers solitary or in capitula (heads); bracts leaf-like, scarious or membranous, persistent; bracteoles sometimes present. Flowers small, regular, hermaphrodite; rarely the plants dioecious. Perianth-segments 6, in two series, subequal, glumaceous, green or brown, usually membranous at the edges. Stamens 6, opposite and shorter than the perianth-segments, the 3 inner sometimes absent; filaments linear or triangular; anthers basifixed, 2-thecous, introrse, dehiscing longitudinally. Carpels 3, joined; ovary superior, 1-or 3-locular; style rarely absent; stigmas 3. Fruit a loculicidal capsule. Seeds 3 or many, ovoid to obovoid, apex sometimes apiculate, base sometimes tailed
Herbs, perennial, occasionally annual, usually rhizomatous, orsometimes cespitose. Culms round or flat. Leaves mostly basal; sheath margins fused or overlapping, often with 2 earlike extensions (auricles) at blade junction; blade flat or round, glabrous or margins hairy. Inflorescences of headlike clusters or single flowers variously arranged; bracts subtending inflorescence 1 or more2, mostly leaflike; bracts subtending inflorescence branches 1--2, reduced; bracteoles subtending solitary flower 0--2, translucent, reduced. Flowers usually bisexual, radially symmetric; sepals and petals similar, persistent, green to brown or purplish black; stamens usually 3 or 6; anthers persistent, linear; pistils 1; ovaries superior, locules 1 or 3, placentas 1 and basal or 3 and axile or parietal; stigmas generally longer than styles. Fruits capsules, loculicidal. Seeds 3--many, often with white appendages on 1 or both ends.
Leaves mostly in a basal tuft, grass-like, linear or filiform, sheathing at the base or reduced to a sheath, sheaths open or closed
Stamens 6 or 3, free; anthers 2-celled, basifixed, opening lengthwise; pollen in tetrads
Seeds sometimes tailed, with a small straight embryo in the middle of endosperm
Perianth-segments 6, in 2 whorls, or rarely only 3, usually glumaceous
Ovary superior, 1-celled or 3-celled; styles and stigmas 1 or 3
Flowers hermaphrodite or unisexual, usually very small
Ovules ascending or parietal, 3 or more
Perennial or annual herbs
Fruit a dry capsule
Life form
Growth form herb
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Sexuality hermaphrodite
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Root system fibrous-root rhizome
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Hardiness (USDA) 7-11

Usage

Juncaceae is of little economic importance in the commonly accepted sense, but the family is an ecologically important and major component in aquatic and other wetland vegetation, providing habitat and nesting material for fauna, substrate stability and other environmental values. Juncus species have been used to help slow water flows, prevent erosion, trap sediments and improve water quality (e.g. contribute to water filtration in constructed wetlands). El-Shamy et al. (2012) cite various sources that give examples of traditional uses of Juncus for medicinal purposes. Some examples include the use of pith from Juncus effusus in traditional Chinese medicine as an antipyretic and also as sedative agent in Japan. Other examples include the use of Juncus seeds as a treatment for diarrhoea, and an infusion of fruits of J. acutus mixed with barley grains to treat the common cold. Split rushes (i.e. Juncus effusus, J. inflexus, J. squarrosus) have been used for making baskets and chair bottoms. A number of species are sometimes available in nurseries, e.g. Juncus effusus (Common Rush, Soft Rush), with several cultivars. Many native species are offered at native plant nurseries. Luzula is fairly rarely cultivated but some species, such as L. nivea (L.) DC. (Snow-white Woodrush) from Europe and L. ulophylla (Buch.) Ckn (Dwarf Woodrush) from New Zealand, are quite ornamental and occasionally offered (HortFlora).
Uses medicinal ornamental wood
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Cultivation

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Images

Juncaceae unspecified picture

Distribution

Juncaceae world distribution map, present in Australia and China

Identifiers

LSID urn:lsid:ipni.org:names:77126749-1
WFO ID wfo-7000000310
COL ID BJG
BDTFX ID 101074
INPN ID 187471
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Juncaceae

Lower taxons

Juncus Luzula Patosia Marsippospermum Oxychloe Distichia Rostkovia Oreojuncus