Luzula Dc.

Woodrush (en), Luzule (fr)

Genus

Angiosperms > Poales > Juncaceae

Characteristics

Perennial herbs, usually tufted and stoloniferous. Leaves for the greater part crowded at the base of an erect or ascending stem, grass-like, linear from a sheathing base, tapering towards the often thickened or shortly acute apex, frequently fringed with long, very fine white hairs. Flowers either solitary or subsolitary and loosely paniculate or crowded into dense clusters; the latter often forming a raceme or panicle which may be contracted into a dense spikelike inflorescence; each flower subtended by a scarious bract and enclosed at the base by 1-2 scarious short bracteoles. Tepals 6, free, glumaceous, equal or slightly unequal, ovate-lanceolate, acute or partly obtuse. Stamens 6, singly before the tepals; filaments thin; anthers erect, oblong or linear. Ovary sessile, 1-celled; ovules 3, erect from a very short basal placenta; style 1, deciduous; arms 3, usually slender. Fruit ellipsoid or obovoid, often crowned by the style-base, 3-valved. Seeds 3 or fewer, oblong, faintly reticulate-ribbed, often with a basal or apical elaiosome; embryo small, near the hilum; albumen subfarinaceous.
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Herbs, perennial, usually cespitose, often with short, mostly vertical to running rhizomes and/or (less commonly) stolons. Culms round. Cataphylls absent. Leaves: sheaths closed, without auricles at throat (junction with blade), usually pilose; blade flat or channeled, never septate, margins with long, soft, multicellular hairs, apex often thickened (callous), veins commonly indistinct. Inflorescences terminal; flowers inserted individually or in dense clusters (glomerules) variously arranged; bracts subtending inflorescence (proximal inflorescence bracts) 2, mostly leaflike; bracts subtending inflorescence branches 1--2, reduced; bracteoles subtending flowers 2--3. Flowers: tepals 6, in 2 whorls; stamens 6. Capsules 1-locular, generally globose; beak often formed by persistent style base. Seeds 3, globose to ovoid, base often with tuft of fibrous hairs (vestige of funiculus); nutritive appendage from outer seed coat (caruncle) often present, white, barely visible to ± equaling seed body. x = 6.
Infl. a terminal cyme, congested to a single head, or much-branched, branchlets unequal in length with fls clustered at their tips; subtending bracts lf-like. Fls hermaphrodite, each enclosed at the base by 2 papery bractlets and subtended by a scarious bract; bracts and bractlets silver or light brown, membr., ovate, with ± lacerate, ± villous margins. Stamens 3, or 6. Ovary 1-locular. Seeds 3, with a white basal tail (caruncle), testa minutely striated in a reticulate pattern. Perennial grasslike herbs, tufted or stoloniferous, or cushion-forming. Stems glab., terete, erect, often with 1–2 cauline lvs. Lvs flat or channelled, almost glab., to densely villous along the margin, basal sheath without auricles. A cosmopolitan genus of c. 80 spp. mainly in extra-tropical regions; in the tropics at high altitudes only. 10 N.Z. spp. are endemic while L. traversii is recorded also from Mexico and South America. Luzula DC. nom. cons.: type sp., L. campestris (L.) DC.
Perennial grass-like herbs, tufted, or stoloniferous or rhizomatous, or cushion-forming. Stems glabrous, terete, erect, often with 1-2 cauline leaves. Leaves flat or channelled, almost glabrous to densely villous along margin, sheaths without auricles. Inflorescence a terminal cyme, congested to a single head, or much-branched, branches unequal with flowers clustered at tips; subtending bracts leaf-like. Flowers hermaphrodite; bracts and bractlets silver or light brown, membranous, ovate, margins ± lacerate, ± villous. Tepals 6, usually equal in length, with or without membranous margins. Stamens 3, or 6. Ovary 1-locular. Seeds 3, with a white basal caruncle (tail). A cosmopolitan genus of c.80 spp., mainly in extra-tropical regions; in the tropics at high altitudes only. Native spp.12, adventive 4.
Herbs perennial, usually tufted. Rhizome short. Stems usually terete. Leaves mostly basal; leaf sheath closed, auricles absent; leaf blade lanceolate to linear, flat, usually channeled, margin long white ciliate. Inflorescences cymose, umbellate, umbel-like, corymbose, or paniculate, sometimes condensed into heads. Flowers often solitary, subtended by a scarious bract and enclosed at base by 2 short bracteoles; bracteoles usually lacerate or denticulate at margin. Perianth segments 6. Stamens 6, usually shorter than perianth; filaments thin; anthers oblong to linear. Ovary 1-loculed; ovules 3, erect from a basal, very short placenta. Style short. Capsule 3-valved. Seeds 3, oblong, indistinctly reticulate, often with a basal or apical appendage (caruncle).
Perennial, rarely annual herbs, tufted, rhizomatous or stoloniferous. Leaves grass-like, flat, basal and cauline, with long hairs on the margins, sheathing below, sometimes reduced to cataphylls; sheaths always closed. Inflorescence compound, panicled or subspicate, bracteate; bracts hairy on the margins; bracteoles present, placed immediately below and clasping the flower. Flowers hermaphrodite. Perianth-segments free, ovate or lanceolate, persistent, margins entire or sometimes lacerated. Stamens 3 or 6, attached at the base of the segments; filaments filiform; anthers oblong or linear. Ovary sessile; style filiform; stigmas filiform. Capsule 1-celled; seeds 3, basal, globose or ovoid, often tailed.
Fls generally prophyllate, as in spp. of Juncus; perianth as in Juncus; ovary and loculicidal capsule unilocular; ovules and seeds 3, basal; infl varying from open, lax, and subumbellately paniculiform to composed of several compact spikes or heads, these sometimes clustered to form a secondary spike or head; lvs with closed sheath and flat, grass-like blade provided (at least when young) with some long, loose hairs along the margins, especially at the throat. 75, mainly in temp. and cool parts of the N. Hemisphere.Spp. 3–7 belong to a circumboreal pillar-complex, perhaps better treated with fewer binomials and more trinomials, but the taxonomy and nomenclature not yet clear.
Grass-like perennials, tufted or stoloniferous. Leaves flat, fringed with few or many long hairs. Inflorescence terminal, cymose, congested into a head or unequally branched, with leaf-like subtending bracts. Flowers with subtending papery or scarious bracts. Stamens 3 or 6. Ovary 1-locular with 3 basal ovules. Fruit a capsule. Seeds shining, usually with a conspicuous light-coloured caruncle.
Stoloniferous, tufted perennials; stems scapiform, foliate chiefly in a basal rosette; leaves grass-like, the basal sheaths closed; inflorescence paniculate, bearing very many small, glumaceous, perfect flowers.
Life form perennial
Growth form herb
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Sexuality hermaphrodite
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Root system rhizome
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Environment

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Hardiness (USDA) 4-10

Usage

Some species, including cultivars, are planted as ornamental 'grasses', e.g. natives of Europe Luzula nivea (Nathh.) DC. (Snow-white Woodrush, Snowy Woodrush, Lesser Woodrush) and L. sylvatica (Huds.) Gaudin (Greater Woodrush).
Uses ornamental wood
Edible -
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Cultivation

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Images

Luzula unspecified picture

Distribution

Luzula world distribution map, present in Australia, China, Malaysia, New Zealand, Panama, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:331233-2
WFO ID wfo-4000022464
COL ID 5HGS
BDTFX ID 86741
INPN ID 194322
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Luzula Juncoides Juncodes

Lower taxons

Luzula ovata Luzula ulei Luzula acuminata Luzula atlantica Luzula atrata Luzula capitata Luzula comosa Luzula confusa Luzula desvauxii Luzula echinata Luzula ecuadoriensis Luzula elegans Luzula excelsa Luzula groenlandica Luzula hawaiiensis Luzula hitchcockii Luzula kjellmaniana Luzula masafuerana Luzula mendocina Luzula meridionalis Luzula oligantha Luzula orestera Luzula ostenii Luzula pallescens Luzula papuana Luzula parviflora Luzula pediformis Luzula rufa Luzula seubertii Luzula spicata Luzula subcapitata Luzula subcongesta Luzula tristachya Luzula wahlenbergii Luzula x hermannii-muelleri Luzula levieri Luzula hasleri Luzula johannis-principis Luzula pfaffii Luzula bornmuelleriana Luzula sichuanensis Luzula wettsteinii Luzula borreri Luzula x romanica Luzula divulgatiformis Luzula exspectata Luzula brevispicata Luzula abchasica Luzula abyssinica Luzula acutifolia Luzula africana Luzula alpestris Luzula alpina Luzula antarctica Luzula arcuata Luzula banksiana Luzula brachyphylla Luzula bulbosa Luzula caespitosa Luzula calabra Luzula campestris Luzula denticulata Luzula divaricata Luzula flaccida Luzula glabrata Luzula indica Luzula jimboi Luzula johnstonii Luzula kobayasii Luzula luzulina Luzula mannii Luzula modesta Luzula multiflora Luzula nodulosa Luzula novae-cambriae Luzula philippinensis Luzula picta Luzula pindica Luzula piperi Luzula plumosa Luzula poimena Luzula purpureosplendens Luzula rufescens Luzula stenophylla Luzula subsessilis Luzula sudetica Luzula sylvatica Luzula taiwaniana Luzula taurica Luzula traversii Luzula vulcanica Luzula x gayana Luzula x danica Luzula x heddae Luzula alopecurus Luzula canariensis Luzula longiflora Luzula lutescens Luzula pumila Luzula x favratii Luzula x somedana Luzula melanocarpa Luzula winderiae Luzula havaiiensis Luzula cascadensis Luzula macrantha Luzula pedemontana Luzula nivalis Luzula vinesii Luzula caricina Luzula chilensis Luzula effusa Luzula leiboldii Luzula peruviana Luzula australasica Luzula colensoi Luzula crinita Luzula densiflora Luzula nipponica Luzula ruiz-lealii Luzula ulophylla Luzula alopecuroides Luzula x hybrida Luzula ignivoma Luzula bomiensis Luzula celata Luzula crenulata Luzula decipiens Luzula divulgata Luzula fallax Luzula formosana Luzula leptophylla Luzula x bogdanii Luzula x media Luzula inaequalis Luzula gigantea Luzula lactea Luzula alpinopilosa Luzula lutea Luzula forsteri Luzula luzuloides Luzula racemosa Luzula nivea Luzula congesta Luzula pilosa