Juncus bufonius L.

Toad rush (en), Jonc des crapauds (fr)

Species

Angiosperms > Poales > Juncaceae > Juncus

Characteristics

Annual to 3 dm, the stems terete, sometimes branched; lvs basal and usually also cauline, capillary, 0.5–10 cm × 0.2–1.1 mm, convex on the lower surface, flat or channeled on the upper; fls often cleistogamous, scattered singly along the upper part (often more than half) of the stem, short-pedicellate or subsessile in the axils of hyaline-scarious or partly leafy bracts, each one also closely subtended by a pair of broad, hyaline-scarious bracteoles 1–2.5 mm; tep slender, with greenish midstrip and hyaline margins, mostly 3–6 mm, the outer a little the longer; anthers 6, 0.3–1 mm, up to as long as the filaments; style very short, fr imperfectly trilocular; seeds 0.3–0.6 mm, apiculate; 2n=26–120, a small polyploid pillar-complex. Moist or vernally wet or moist places; nearly cosmopolitan, and throughout our range. Most of our plants are polyploid and have acuminate pet slightly longer than the fr. A ± diploid halophytic phase, found in salt-marshes along the coast from Lab. to Mass., and widespread in Europe, differs morphologically from typical, widespread J. bufonius in its consistently small size (rarely to 15 cm) and obtuse or mucronate pet equaling or slightly shorter than the fr. This phase has been segregated as var. halophilus Fernald or J. ambiguus Guss.
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Very variable, pale-coloured annual, usually 10-30 cm long, rarely longer. Stems tufted, all fertile, erect, erecto-patent or decumbent, slender, terete, soft, often branched, leafy. Leaf-sheaths rather long, not auricled. Leaves linear, very narrow, channelled down the face, the lowmost up to 16 cm long, the higher shorter. Inflorescence terminal, branched; branches erect or obliquely erect, rather straight. Flowers either solitary and remote or in more or less remote (sometimes almost crowded) clusters of 2-6; each flower at the base with a bract and 2 bracteoles; flowers nearly sessile, pale-coloured, very variable as to size, 3-7 mm long. Bracts and bracteoles thinly membranous, ovate, acute. Tepals narrowly ovate-lanceolate, very acute, the 3 inner often much smaller than the outer. Stamens 6, in the topmost flowers often 3. Style distinct; stigmas straight. Capsule oblong, trigonous, usually shorter than the perianth, 3-5 mm long; placentas at length often detached from the valves, either free or cohering into a central column. Seeds numerous, ellipsoid, tailless, brown, extremely faintly ribbed, ⅓-½ mm long.
Plants annual, 4--20(--30) cm tall. Stems usually many, erect or ascending, usually weak, base often reddish brown. Leaves basal and cauline; leaf sheath often swollen basally, margin hyaline; leaf blade flat, 1--13 cm × ca. 1 mm, margin often slightly convolute. Inflorescences usually lax, 1/4--4/5 × plant height; branches few, ascending, somewhat arcuate, usually weak, few flowered. Flowers subterminal, remote, appressed to subappressed; involucral bract leaflike. Bracteoles 2 or 3, 1.3--2.5 × 1.2--2.2 mm. Perianth segments lanceolate; outer segments 4.25--5.5 × 1--1.8 mm, center green, margin broadly white membranous, apex sharply pointed; inner segments 4--5 mm, often mostly membranous, apex acute. Stamens 6, 1/3--1/2 as long as perianth; filaments 0.8--1.2 mm; anthers 0.5--0.8 mm. Style absent; stigmas ca. 0.5 mm. Capsule ovoid to subellipsoid, 3--4(--5) mm, 3-loculed, apex slightly obtuse. Seeds ellipsoid, 0.4--0.6 mm, striate, base and apex usually apiculate. Fl. May--Jul, fr. Jun--Sep. 2 n = 100, 110.
Herbs, annual, cespitose, 0.5--4 dm. Culms 1--many, occasionally becoming decumbent. Cataphylls 0--2. Leaves basal and cauline; auricles rudimentary or absent; blade flat, 3--13 cm x 0.3--1.1 mm. Inflorescences loose and diffuse or less often compact, usually at least ½1/2 total height of plant; primary bract shorter than inflorescence. Flowers: bracteoles 2; tepals greenish, lanceolate, 3.8--7(--8.5) mm; inner series slightly shorter, apex sometimes obtuse; stamens 3--6, filaments (0.7--)1--1.8 mm, anthers 0.3--0.8 mm; style 0.1--0.2 mm. Capsules tan to reddish brown, 3-locular, ellipsoid to narrowly so, slightly truncate, 2.7--4 x 1--1.5 mm, sometimes exceeding inner tepals but usually not outer series. Seeds yellowish, widely ellipsoid to ovoid, 0.26--0.49, not tailed. 2n = 27--37, 58--81, 108--115.
Annual, rarely longer lived, tufted, light green or, in dry situations, often red-tinged, extremely variable, (2.5)-10-20-(40) cm high, whole plant easily pulled out of ground, roots fibrous. Stems simple or branched near base, branching above to form inflorescence. Leaves few, mostly basal, occasionally cauline, ± setaceous, ± channelled, auricles absent. Inflorescence highly variable in size and shape, often ⅔ length of whole plant. Flowers sessile, borne singly or in 2-7-flowered clusters. Tepals (3)-4-6-(9) mm long, outer tepals longer-acuminate than inner. Stamens (3)-6.Capsule (2.5)-3-4 mm long, < tepals, oblong, blunt, green to bright red-brown.
Annuals, tufted, soft, 30-300 mm tall. Leaves few, radical and cauline, narrowly linear-acuminate, with an orange, loosely sheathing base. Flowers in leafy panicles or lateral secund spikes, sessile, solitary; bracts 3, hyaline. Tepals c. 5 mm long, green with a hyaline border, outer aristate, inner shorter, acute. Stamens 6(-3), short; anthers about as long as filaments. Ovary 3-locular; style and stigmas about as long as perianth. Capsule included, ellipsoid, obtuse; seed obovoid, c. 0.4 mm long, base rounded, apex truncate, ribbed, pale brown, becoming mucilaginous when wet. Flowering in spring.
Annual herb, tufted, up to 300 mm high. Leaves few, radical and cauline, narrowly linear-acuminate. Flowers in much-branched, exserted panicles (anthelae), or lateral second spikes, sessile, solitary; bracts 3, hyaline. Perianth segments hard, green with a hyaline border, glumaceous. Stamens: anthers ± as long as filaments. Ovary with 3 ± separate locules; ovules biseriate, many; style terete with 3 stigmatic arms, ± as long as perianth. Capsule loculicidal, included, ellipsoid, obtuse. Seeds many, 0.4 mm long, apex truncate. Flowering time Jan.-Dec.
Soft annual, up to 300 mm high. Stems leafy. Leaves forming loose basal rosettes; blade flat, linear, base not persisting; cataphylls absent. Flowers: solitary or in clusters of 2-5, in leafy panicles, each flower with a bract and 2 bracteoles; perianth green; Oct.-Jan.
Soft annual, 15-30 cm. Leaves linear. Flowers in leafy, branched spikes, green.
Life form annual
Growth form herb
Growth support free-standing
Foliage retention
Sexuality hermaphrodite
Pollination
Spread epizoochory
Mature width (meter) -
Mature height (meter) 0.13 - 0.3
Root system fibrous-root
Rooting depth (meter) 0.2
Root diameter (meter) 0.1
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

Permanently or temporarily humid localities, often gregarious. In Luzon along bridle-trails in mossy forests at 1900 m.
Light 7-8
Soil humidity 4-7
Soil texture 1-3
Soil acidity 2-7
Soil nutriment -
Hardiness (USDA) 5-10

Usage

Uses animal food environmental use medicinal
Edible -
Therapeutic use Emetic (unspecified), Dermatological Aid (unspecified), Strengthener (unspecified), Diarrhea (unspecified), Exanthema (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by divisions or seedlings.
Mode divisions seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Juncus bufonius habit picture by Pierre LEON (cc-by-sa)
Juncus bufonius habit picture by Yoan MARTIN (cc-by-sa)
Juncus bufonius habit picture by Yoan MARTIN (cc-by-sa)

Leaf

Juncus bufonius leaf picture by Pierre LEON (cc-by-sa)
Juncus bufonius leaf picture by P C (cc-by-sa)
Juncus bufonius leaf picture by Gerrit Busser (cc-by-sa)

Flower

Juncus bufonius flower picture by Pierre LEON (cc-by-sa)
Juncus bufonius flower picture by Jacques Maréchal (cc-by-sa)
Juncus bufonius flower picture by Jiří Hedrich (cc-by-sa)

Distribution

Juncus bufonius world distribution map, present in Afghanistan, Anguilla, Albania, Andorra, Argentina, Armenia, Austria, Azerbaijan, Belgium, Bangladesh, Bulgaria, Bosnia and Herzegovina, Belarus, Bolivia (Plurinational State of), Brazil, Bhutan, Canada, Switzerland, Chile, China, Colombia, Costa Rica, Czech Republic, Germany, Denmark, Algeria, Ecuador, Egypt, Eritrea, Spain, Estonia, Ethiopia, Finland, France, United Kingdom of Great Britain and Northern Ireland, Georgia, Gibraltar, Guadeloupe, Greece, Guatemala, Croatia, Hungary, India, Iran (Islamic Republic of), Iraq, Iceland, Israel, Italy, Jamaica, Japan, Kazakhstan, Kenya, Kyrgyzstan, Lebanon, Liberia, Libya, Liechtenstein, Sri Lanka, Lithuania, Luxembourg, Latvia, Morocco, Mexico, North Macedonia, Malta, Myanmar, Montenegro, Mongolia, Mauritius, Malaysia, Namibia, Netherlands, Norway, Nepal, New Zealand, Pakistan, Peru, Philippines, Poland, Korea (Democratic People's Republic of), Portugal, Réunion, Romania, Russian Federation, Rwanda, Saudi Arabia, Sudan, Saint Helena, Ascension and Tristan da Cunha, Saint Pierre and Miquelon, Serbia, Slovakia, Slovenia, Sweden, Syrian Arab Republic, Chad, Thailand, Tajikistan, Turkmenistan, Tunisia, Turkey, Taiwan, Province of China, Uganda, Ukraine, Uruguay, United States of America, Uzbekistan, Venezuela (Bolivarian Republic of), Viet Nam, and South Africa

Conservation status

Juncus bufonius threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:442769-1
WFO ID wfo-0000776030
COL ID 3QTK3
BDTFX ID 36441
INPN ID 104144
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Juncus bufonius f. filiformis Juncus bufonius f. giganteus Cyperus bufonius Juncus prolifer Tenageia bufonia Juncus aletaiensis Juncus fasciatus Juncus istriacus Juncus creticus Juncus pumilus Juncus cespifolius Juncus bilineatus Juncus bufonius f. alpinus Juncus bufonius f. bracteatus Juncus bufonius f. divaricatus Juncus divaricatus Juncus erythropodus Juncus bufonius f. bufonius Juncus bufonum Juncus plebeius Juncus juzepczukii Juncus leptocladus Schoenus ferrugineus Juncus congestus Juncus bufonius var. capillaris Juncus bufonius var. compactus Juncus bufonius var. congestus Juncus bufonius var. dolichophyllus Juncus bufonius var. filiformis Juncus bufonius var. gramineus Juncus bufonius var. grandiflorus Juncus bufonius subsp. juzepczukii Juncus bufonius var. juzepczukii Juncus bufonius var. laxus Juncus bufonius var. procerior Juncus bufonius var. pumilio Juncus bufonius var. subauriculatus Juncus bufonius var. suffocatus Juncus bufonius var. susianus Juncus bufonius var. tuberiferus Juncus bufonius var. typicus Juncus bufonius var. gracilis Juncus bufonius var. alpinus Juncus bufonius var. bufonius Juncus bufonius f. scoparius Juncus bufonius