Leea D.Royen

Trees or shrubs, sometimes scandent, without tendrils. Leaves alternate, usually 1–3-pinnate; leaflets opposite, crenate to serrate, glandular; petiole and stipules fused, expanded, sheathing around stem apex. Inflorescence cymose, leaf-opposed, bracteate, often condensed. Flowers actinomorphic, bisexual. Calyx campanulate, 4-or 5-lobed. Corolla hypogynous, 4-or 5-lobed, valvate, tubular at base. Disc absent. Epipetalous tube present, consisting of a lower ring extending down towards ovary, and an upper corona which is 4-or 5-lobed. Stamens epipetalous, from base of corona; filaments flattened, passing over sinuses between lobes; anthers coherent in a ring inside corona, dehiscing by longitudinal slits. Ovary superior, discoid, usually 4–6-locular, each with 1 ovule. Style short, entire. Fruit a depressed-globose berry with up to 6 seeds. Seeds without a distinct beak; endosperm present.