Herbs annual. Trichomes unicellular, eglandular, simple, mixed with much fewer and smaller, glandular, multicellular ones with multiseriate stalk. Stems erect, simple or branched above. Basal leaves petiolate, not rosulate, simple, pinnatifid, dentate, or rarely entire, often withered by flowering. Cauline leaves petiolate or subsessile, entire or dentate. Racemes ebracteate, elongated in fruit. Fruiting pedicels slender, ascending, articulate at rachis. Sepals oblong, erect, base of lateral pair not saccate. Petals white or purplish, longer than sepals; blade narrowly spatulate or oblong, apex obtuse; claw subequaling sepals. Stamens 6, slightly tetradynamous; filaments flattened at base; anthers ovate, obtuse at apex. Nectar glands 6; lateral semiannular; median very small, 1 facing each median filament. Ovules 2 per ovary, subapical. Fruit indehiscent, nutletlike silicles, globose, terete, sessile; valves thickened, woody, prominently veined, vesicular papillate; replum strongly expanded laterally; septum complete, thickened, opaque, veinless; style prominent and to 3.5 mm, slender, cylindric, articulated at base, straight, caducous, glabrous; stigma capitate, entire. Seeds wingless, orbicular, strongly flattened; seed coat reticulate, not mucilaginous when wetted; cotyledons accumbent.