Lolium temulentum L.

Darnel ryegrass (en), Ivraie enivrante (fr), Ray-grass commun (fr)

Species

Angiosperms > Poales > Poaceae > Lolium

Characteristics

Annual. Culms 32–100 cm high, often rooting from lower nodes; mid-culm nodes often reddish. Leaves: basal sheaths not pigmented; auricles usually present; blade convolute in bud, 13.5–28 cm long, 4–10 mm wide. Spike 8.5–29.5 cm long, with 5–22 (–30) spikelets; axis rigid, 1.3–3 mm wide at lowest node, with angles scabrous. Spikelets to 1.5 spikelet-lengths apart, 10–27 mm long, 3–8 mm wide, with 3–9 (–11) bisexual florets; rachilla segments 1.7–3.8 mm long (longer at base). Glume slightly shorter than to slightly exceeding remainder of spikelet, conspicuously longer than basal lemma; upper glume narrowly oblong, 10–24 mm long, 5–9-nerved. Lemma laterally compressed, (4.5–) 6–8.5 mm long, awnless or awned, chartaceous to indurated, 5 or 7-nerved, rarely becoming pigmented toward margin at apex; awn to 17 mm long. Caryopsis very swollen at maturity, oblong, longitudinally grooved, (3.8–) 4–7 mm long, (1.1–) 1.6–3 mm wide.
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Stiff, stout, erect annuals, 35-100 cm; branching intravaginal. Leaf-sheath glabrous, light brown. Auricles to 1 mm, spreading, or 0. Ligule 0.8-1.5 mm, truncate. Leaf-blade 10-22 cm × 3-4.5-(6) mm, flat, abaxially glabrous, adaxially scaberulous or rarely smooth; margins smooth below, scabrid near tapered acute tip. Culm 20-60 cm, internodes closely scabrid above or smooth. Spike 10-20-(34) cm, erect; rachis 1-2 mm wide, smooth or finely scabrid on convex side, margins scaberulous. Spikelets 10-30 mm, 5-12-flowered, green; florets turgid at maturity. Upper glume 10-22 mm, < to > spikelet, 7-9-nerved, rigid, oblong-lanceolate, obtuse or subacute, smooth or scabrid. Lemma 5.5-9.5 mm, elliptic to ovate, smooth below, becoming hard at maturity, apex hyaline, scabrid, bifid, awned from sinus; awn fine, strongly scabrid, 1-13 mm, or 0. Palea = lemma. Anthers 2-3 mm. Caryopsis 5-6 × 2-2.5 mm.
Annual. Culms tufted, erect or decumbent, slender to moderately robust, 20–120 cm tall, 3–5-noded. Leaf blades flat, thin, 10–25 cm × 4–10 mm, smooth or scabridulous on abaxial surface, margins scabrid, young blades rolled; auricles present or absent; ligule 0.5–2.5 mm, obtuse to truncate. Raceme stiff, straight, 10–30 cm; rachis thick, smooth or scabridulous, spikelets about their own length apart. Spikelets turgid, 0.8–2.5 cm, florets 4–10, rachilla internodes 1–1.5 mm, smooth, glabrous; glume linear-oblong, rigid, as long as spikelet, often exceeding florets, 5–9-veined, margins narrowly membranous, apex obtuse; lemmas elliptic to ovate, turgid at maturity, 5.2–8.5 mm, apex obtuse; awn usually present, stiff, scabrid; palea ciliolate along keels. Caryopsis very plump, length 2–3 times width, 4–7 mm. Fl. and fr. May–Aug. 2n = 14.
Robust annual, 400-900 mm high; culms solitary or tufted. Leaf blade 150-300 x 3-7 mm; ligule an unfringed membrane. Inflorescence a spike, spikelets sunk and lying edgeways in partial cavities on alternate sides of central axis. Spikelet 8-28 x 3-8 mm; lower glume usually suppressed, except in terminal spikelet; upper glume ascending or spreading, shorter than spikelet. Florets 5-9; lemma elliptical to ovate, < 3x as long as wide, very turgid at maturity, awn absent or up to 20 mm long; palea as long as lemma, 2-keeled; anther 2.1-3.2 mm long. Flowering time Sept.-Feb. Caryopsis when mature 2-3x as long as wide.
Annual; culms solitary or few together, 4–10 dm, scaberulous; lvs usually glabrous beneath, scabrous above, 3–10 mm wide; spike 1–2(–4) dm with scabrous margins; spikelets 4–8-fld; glume firm, straight, strongly veined, equaling or surpassing the uppermost lemma, (12–)15–25 mm; lemmas obtuse, with an awn to 17 mm, or sometimes awnless, the lower lemmas in the spikelet 5–8.5 mm; paleas shorter than the lemmas, grain 2–3 times as long as thick; 2n=14. Probably native to the Mediterranean region, now widespread as a weed and occasionally found throughout most of our range.
Stiff annual.. Culms 30–90 cm. high, tufted or solitary, erect or geniculate at the base, slender to moderately stout.. Leaf-blades 6–40 cm. long and 3–13 mm. wide, flat, with short spreading auricles at the base.. Spikes erect, 10–30 cm. long and 5–12 mm. wide, rigid, green, the spikelets about their own length or more apart.. Spikelets oblong, 12–26 mm. long, 4–10-flowered; upper glume usually reaching to or exceeding uppermost lemma, rigid, 7–9-nerved, obtuse; lemmas elliptic to ovate, 6–8 mm. long, smooth, obtuse, awned or awnless.
Robust annual 400-900 mm high; culms solitary or tufted. Leaf blade 150-300 x 3-7 mm. Spikelet 8-28 x 3-8 mm, 5-9-flowered; lower glume usually suppressed, except in terminal spikelets; upper glume ascending or spreading, 3/4 to 1 1/2 x spikelet length, not concealing the spikelet; lemma elliptical to ovate, less than 3 x as long as wide, very turgid at maturity, awn absent or to 20 mm long; palea as long as lemma, 2-keeled; anther 2.1-3.2 mm long; mature caryopsis 2-3 times as long as wide.
Annual to 1 m tall. Leaves linear; ligule an unfringed membrane. Inflorescence a simple, true spike; 100-300 mm long. Spikelets 8-28 mm long, laterally compressed, glabrous, awnless, sunk and lying edgeways in partial cavities on main axis, separated by ± own length, one edge against rhachis, green, many-flowered; glumes usually 1, with 2 only in terminal spikelets, as long as spikelet; lemmas rounded, swollen at maturity, awnless or with awn up to 20 mm long.
Annual; up to 0.9 m high; robust. Culms solitary or tufted. Leaf blades 150-300 x 3-7 mm. Flowers: panicle a single; flat spike; spikelets 8-28 x 3-8 mm; glumes ascending or spreading; not concealing spikelets; 3/4-1 and1/2 x spikelet length; lemmas elliptical to ovate; very turgid at maturity; awn absent or up to 20 mm long; mature caryopsis 2-3 x as long as wide.
Robust annual, culms solitary or tufted, up to 0.9 m high. Leaf blades 150-300 mm long, 3-7 mm wide. Spikelets 8-28 mm long, 3-8 mm wide. Glumes ascending or spreading, not concealing spikelets, 3/4 to 1 1/2 times spikelet length; lemmas elliptical to ovate, very turgid at maturity, awn absent or to 20 mm long. Mature caryopsis 2-3 times as long as wide.
A stiff short annual grass. It usually forms tufts and has erect stems. The leaves are alternate and in two ranks. The leaves are green and the blades are 3-13 mm wide. The flowers are in erect spikes. They are somewhat zigzag between the spikelets. The spikelets are 8-28 mm long.
Leaf-laminae 6-30(-40) x 0·3-1·3 cm., linear, tapering to a fine point, with 2 narrow auricles at the base, usually expanded, firm, scaberulous or smooth except for the margins.
Lemmas 6-8 mm. long, 5-9-nerved, muticous or awned, dorsally rounded, elliptic to ovate, somewhat rigid, smooth; awn (if present) up to 20 mm. long, straight, scabrous.
Spikelets 12-25 x 4-6 mm., 4-10-flowered, elliptic-oblong, oblong or obovate-oblong in outline, moderately compressed about their own length or slightly more apart.
Culms up to 90 cm. tall, 2-4-noded, solitary or tufted, somewhat stout, erect or geniculately ascending, glabrous, smooth or scabrous towards the spike.
Superior glume 7-9-nerved, as long as or longer than the rest of the spikelet, narrowly oblong, obtuse, rigid, smooth or asperulous.
Leaf-sheaths prominently striate, rather tight, dorsally rounded, smooth or scaberulous, glabrous; Ligule c. 2 mm. long.
Spikes 10-30 cm. long, erect, rigid, pale-green; rhachis stout, somewhat waved, dorsally scaberulous or smooth.
Annual to 1 m. Leaves linear. Spikelets in a simple spike, green, 8-28 mm long, lemmas swollen at maturity.
Caryopsis c. 7 mm. long, elliptic-oblong in outline, tightly enclosed by the lemma and palea.
Anthers c. 2·5 mm. long, linear.
A rather robust annual.
Life form annual
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality hermaphrodite
Pollination anemogamy
Spread epizoochory
Mature width (meter) -
Mature height (meter) 0.9 - 1.0
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

It is a temperate plant. It grows in waste and fallow ground. In Argentina it grows below 700 m above sea level.
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Cultivated fields and waste ground in Britain.
Light 7-9
Soil humidity 3-5
Soil texture 3-6
Soil acidity 3-7
Soil nutriment -
Hardiness (USDA) 5-8

Usage

The seeds have been eaten. CAUTION: Sometimes a poisonous fungus (Endoconidium temulentum) affects the grains.
Uses animal food food gene source medicinal poison vertebrate poison
Edible seeds stems
Therapeutic use Wounds and injuries (seed), Anodyne (unspecified), Homeopathy (unspecified), Intoxicant (unspecified), Narcotic (unspecified), Neuralgia (unspecified), Poison (unspecified), Rheumatism (unspecified), Wen (unspecified), Hypnotics and sedatives (unspecified), Exanthema (whole plant), Gout (whole plant), Hypnotics and sedatives (whole plant), Freckles (whole plant), Ulcer (whole plant)
Human toxicity infested plants only (seed), infested plants only (fruit)
Animal toxicity strong toxic (seed), strong toxic (fruit)

Cultivation

Plants are grown from seed.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Lolium temulentum habit picture by Yoan MARTIN (cc-by-sa)

Distribution

Lolium temulentum world distribution map, present in Afghanistan, Åland Islands, Albania, Australia, Austria, Bangladesh, Bulgaria, Belarus, Bolivia (Plurinational State of), Brazil, Canada, Switzerland, China, Cuba, Cyprus, Czech Republic, Germany, Algeria, Ecuador, Egypt, Eritrea, Spain, Ethiopia, Finland, France, Micronesia (Federated States of), Georgia, Greece, Guatemala, Hungary, Indonesia, Iran (Islamic Republic of), Iraq, Italy, Japan, Kazakhstan, Kenya, Kyrgyzstan, Kuwait, Libya, Lesotho, Morocco, Mauritius, Malaysia, Nicaragua, Norway, Nepal, New Zealand, Oman, Pakistan, Panama, Peru, Philippines, Palau, Poland, Korea (Democratic People's Republic of), Portugal, Réunion, Romania, Saudi Arabia, Singapore, Turks and Caicos Islands, Turkmenistan, Tunisia, Tanzania, United Republic of, Ukraine, Uruguay, United States of America, Uzbekistan, Venezuela (Bolivarian Republic of), Yemen, South Africa, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:407516-1
WFO ID wfo-0000878503
COL ID 72MWC
BDTFX ID 39788
INPN ID 106517
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Lolium temulentum f. leve Lolium aegyptiacum Lolium album Lolium arvense Lolium decipiens Lolium giganteum Lolium speciosum Lolium temulentum Festuca temulenta Lolium asperum Lolium gracile Lolium gracile Lolium gussonei Lolium infelix Lolium maximum Lolium remotum f. asperum Lolium pseudolinicola Lolium temulentum f. arvense Loliola pratensis Lolium temulentum f. temulentum Lolium asperum Lolium annuum Bromus temulentus Lolium berteronianum Lolium cuneatum Lolium longiglume Lolium lucidum Lolium maximum Craepalia temulenta Lolium robustum Lolium triticoides Lolium temulentum subsp. arvense Lolium temulentum subsp. cuneatum Lolium temulentum subsp. speciosum Lolium temulentum var. arvense Lolium temulentum var. leptochaeton Lolium temulentum var. macrochaeton Lolium temulentum var. muticum Lolium temulentum var. semiglabrum Lolium temulentum var. speciosum Lolium temulentum f. macrochaeton Lolium temulentum f. muticum Lolium temulentum var. temulentum Lolium temulentum var. speciosum