Trees or shrubs; young growth rusty-tomentose. Leaves alternate, simple or compound, entire or divided, sometimes heteromorphic. Conflorescences axillary or terminal, pseudo-racemose, laterally bearing flower pairs, basitonic or acrotonic, solitary or aggregated. Flowers usually fragrant, bisexual; bracts small, caducous or absent. Perianth zygomorphic, slit on lower side in bud; limb globular, recurved in bud; segments eventually separating. Anthers sessile. Hypogynous glands 3. Style long, bent at 90° to ovary; pollen-presenter a flat, oblique disc. Ovary on a long gynophore, glabrous; ovules many in 2 rows, imbricate upwards. Fruit an ovoid follicle with persistent style, coriaceous, opening flat. Seeds ±triangular, flat, covered with greenish yellow granular material, with a translucent wing, surrounded by marginal raphe.