Shrubs, subshrubs, rarely herbaceous, rarely epiphytic; branches rounded to quadrangular, longitudinally canaliculate, the internodes short to elongated. Leaves cauline, subsessile to petiolate; the petiole often amplexicaul; the lamina obovate, ovate to lanceolate, acute to cuspidate at the apex, usually cuneate at the base, membranous to coriaceous. Inflorescences terminal and axillary, open cymose, paniculate, or rarely racemose, often few-flowered. Flowers 5-merous; calyx campanulate to subtubular, the lobes imbricate or rarely valvate, usually unequal; corolla white, yellow, greenish, or red, salverform, narrow to broadly campanulate, rarely tubular, the lobes usually unequal, usually ovate; stamens adnate at or near the middle of the corolla tube, generally included, the filaments flattened, the anthers versatile, usually ovate, bilobate at the base; pistil often exserted, the ovary fusiform to pyriform, the style filiform, elongate, the stigma bilobate, often deeply so. Capsules ovoid to fusiform, often beaked by the per-sistent style, septicidally 2-valvate; seeds numerous, minute. Pollen grains in monads, radially symmetrical, isopolar, oblate, spheroidal, the amb rounded-triangular, 41-45 x 46-53 microns, 3-colporate to 3-porate, the colpi relatively short with rounded ends, the colpus membrane granulose, the ora lalongate to lolongate, the pores oval to circular, the diameter of apocolpia (apoporia) 35-36 microns; exine 3-4 microns thick; sexine thicker than nexine, reticulate, heterobrochate, the lumina 0.5-10 l in diameter, usually rounded, the muri 1-3 ,u wide, supported by one or usually more rows of bacules, of a pilum-or clava-like shape with rounded to truncate apices; nexine generally granulose (M. browallioides).
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Branched to nearly unbranched shrubs, treelets, or trees 1-10 m high. Leaves sessile or petiolate, 3-60 cm long; camptodromous-veined. Inflorescences terminal, cymose or sometimes a thyrse; bracts and bracteoles leaf-like or scale-like towards inflorescence apex. Flowers pedicellate, 5-merous, slightly zygomorphic; calyx fused for ca. 1/2 of length, leathery to thin-leathery, campanulate, lobes of various shapes, with dorsally thickened glandular area, not keeled or winged, apex of various shapes; corolla bud apex rounded; corolla fused for ca. 3/4 of length, fleshy in most species, funnel-shaped, tubular, or salver-shaped; androecium zygomorphic with stamens and style bent towards bottom of corolla mouth; stamens inserted in corolla tube, of unequal length, base of filaments without appendages or a corona, anthers linear; pollen in monads; pistil with nectary disk around base, stigma bilamellate. Fruit a capsule, with calyx persistent and corolla deciduous; seeds angular, rounded, or flat, testa pattern with concave and/or dome-like cells.