Trees, dioecious or monoecious. Leaves alternate, distichous on the lateral branches; stipules amplexicaul, free; blade entire, pinnately veined. Inflorescences on spurs in the leaf axils, unisexual, discoid-capitate, involucrate. Staminate inflorescences mostly several together, pedunculate; flowers several to numerous, perianth 4-lobed to 4-parted; stamens (3-)4; pistillode absent. Pistillate inflorescences mostly solitary, subsessile or pedunculate; flowers many and free, a few and connate, or one; perianth 4-lobed to 4-fid; ovary adnate to the perianth; stigmas 2, tongue-shaped. Fruiting perianth enlarged, fleshy, orange; fruit adnate to the perianth; seed large, without endosperm, cotyledons thick and equal.