Herbs prostrate or erect, often somewhat viscid. Leaves opposite, simple, petiolate, pinnately-nerved, membranaceous or rugose, generally dentate. Inflores-cences capitula, loosely-flowered and pedunculate, or rarely flowers solitary, the bracts linear-lanceolate to ? ovate or awl-shaped. Flowers sessile or shortly pedi-cellate; bracteoles not apparent; calyx campanulate or funnelform, ? actinomor-phic, with 5 ? equal lobes, lanceolate to deltoid; corolla tubular, bilabiate, the upper lip 2-lobed, the lower lip prominently 3-lobed, the middle lobe saccate; stamens 4, paired, mounted near the top of the corolla throat, decurved into the lower saccate lip, the filaments short, distinct, the connective not developed, the anthers with 2 functional thecae; ovary 4-lobed, the gynobase unlobed, the style bifid near the tip, the branches ? even. Nutlets 4, attached laterally to a columnar extension of the gynobase and ? open along this line at maturity.