Molluginaceae Bartl.

Molluginacées (fr)

Family

Angiosperms > Caryophyllales

Characteristics

Herbs, annual or perennial, or sometimes small shrubs or subshrubs, bisexual or rarely dioecious (outside Australia), usually glabrous but glandular or stellate hairs or papillae sometimes present. Leaves alternate, opposite, in false whorls, or crowded in a basal rosette, simple, entire, often mucronate to aristate at the apex; stipules membranous, often fimbriate-laciniate, but sometimes small, obsolete or absent. Flowers actinomorphic, small, in terminal or seemingly axillary, umbel-, panicle-or raceme-like or ± dichotomously branching cymes, or flowers solitary; perianth simple; tepals (4 or) 5, free or sometimes basally united, with membranous margins and with the upper surface often petaloid, with quincuncial aestivation; staminodes rarely present, sometimes petaloid; nectariferous disc sometimes present. Stamens 3–5, 10, 15 or rarely more; filaments free or rarely connate at the base (outside Australia); anthers dehiscing by longitudinal slits; pollen grains usually tricolpate, sometimes polypantocolpate or polypantoporate; carpels 2–5, connate, or carpel solitary (outside Australia); ovules usually few to many per carpel, but sometimes only one, with axile or seemingly basal placentation; styles 1–5, sometimes ± connate at the base and forming a single 2–5-lobed style; stigmas linear or rarely capitate. Fruit a loculicidal capsule or a nutlet (outside Australia). Seeds usually subreniform–suborbicular in outline; aril usually absent or obsolete, but sometimes prominent.
More
Herbs annual or perennial, subshrubs, or shrubs, rarely dioecious, glabrous or rarely hairy. Stems erect or prostrate. Leaves simple, alternate, rarely opposite, often in a basal rosette or in pseudowhorls on stems, margin entire; stipules absent or membranous. Inflorescences terminal or in seemingly axillary cymes, rarely as a solitary flower. Flowers bisexual, rarely unisexual, actinomorphic, hypogynous, rarely perigynous. Tepals 5, rarely 4, free or connate below into a tube, lobes white or pink to purple, sometimes yellow inside (in Glinus). Petals absent or few to many, white, pink, or purple. Stamens 3-5 or many, arranged in several rings, free or connate at base in bundles; anthers dehiscing by longitudinal slits. Ovary superior, syncarpous (in Gisekia developmentally syncarpous; fruit a deeply lobed schizocarp); carpels 2-5 or many, placentation axile, rarely seemingly basal. Stigmas as many as locules. Ovules 1 to many per locule. Fruit usually a loculicidal capsule or deeply (3-)5-15-lobed mericarps (in Gisekia), rarely breaking into 2 nutlets [in Limeum Linnaeus, not in Flora area]. Seeds with embryo curved around a hard, starchy perisperm.
Herbs [shrubs], annual or perennial, glabrous or pubescent, not or slightly succulent. Leaves alternate, opposite, or whorled, petiolate [or sessile]; blade margins entire; stipules present or absent. Inflorescences axillary or terminal, cymose or umbellate, or flowers solitary. Flowers usually bisexual, sessile or pedicellate. Perianth hypogynous; sepals 4-5, distinct to basally connate; petals absent, or small and of staminodial origin, distinct or basally connate; stamens 2-25, hypogynous, in 1-2 series, sometimes fascicled, distinct or basally connate; filaments filiform; anthers versatile, 4-locular, dehiscence introrse, longitudinal; pollen 3-aperturate; pistils of 1-5 connate carpels; placentation axile or appearing basal in some 1-ovulate carpels; ovules 1-25 per locule; stigmas 1 or 3-5, apical, sessile or borne on styles. Fruits achenes or 3-5-valved loculicidal capsules. Seeds reniform to lenticular, sometimes strophiolate (with hilar appendages); endosperm lacking, starch present; embryo curved.
Stamens 3–?, hypogynous, sometimes in pairs or fascicles, alternate with the perianth-segments when definite in number; filaments often enlarged below; anthers 2-locular, opening by longitudinal slits
Ovary superior, 2–5-carpellate, apocarpous (in Gisekia) or syncarpous; loculi and stigmas as many as carpels; ovules 1, few or many per loculus; placentation axile or basal
Succulent or subsucculent annual or perennial herbs or subshrubs, glabrous or covered with simple or stellate hairs (in Glinus lotoides)
Stamens hypogynous or slightly perigynous, definite or many; filaments free to connate at the base; anthers 2-celled, opening lengthwise
Ovary syncarpous (except Gisekia), usually several-celled; styles or stigmas as many as the cells; ovules curved or inverted
Fruit dry, dehiscing into the cells or by a transverse slit, rarely indehiscent, calyx usually persistent
Inflorescences solitary, cymose, loosely dichasial to umbelliform or glomerate, axillary or terminal
Seeds usually subreniform or triangular in outline, sometimes strophiolate; embryo usually curved
Leaves simple, alternate, opposite or verticillate, sometimes crowded, with or without stipules
Flowers more or less actinomorphic, hermaphrodite, rarely unisexual (in Gisekia africana)
Flowers actinomorphic, hermaphrodite, solitary or cymose, usually small and inconspicuous
Leaves alternate or opposite; stipules none or very small and soon falling off
Perianth-segments 5, free, imbricate, herbaceous, persistent
Fruit of achenes, mericarps or loculicidal capsules
Sepals free or united at the base only, imbricate
Staminodes often present, sometimes petaloid
Seeds with endosperm, the embryo curved
Petals small or absent
Herbs
Life form
Growth form herb
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Sexuality hermaphrodite
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Environment

Mostly found in seasonally damp depressions and drainage lines; a few species in arid, rocky habitats.
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Usage

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Cultivation

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Images

Molluginaceae unspecified picture

Distribution

Molluginaceae world distribution map, present in Australia and China

Identifiers

LSID urn:lsid:ipni.org:names:77126662-1
WFO ID wfo-7000000388
COL ID 624P2
BDTFX ID 101034
INPN ID 187281
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Molluginaceae

Lower taxons

Polpoda Mollugo Telephium Glinus Pharnaceum Glischrothamnus Suessenguthiella Adenogramma Coelanthum Paramollugo Hypertelis Psammotropha