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Staminate inflorescences branched; flowers in spike-like to globose glomerules; tepals 3–4; stamens 3–4.
Fruiting perianth enlarged, fleshy, yellow to orange-red; fruit adnate to the perianth, endocarp woody.
Lamina basally attached, palmately incised (sometimes entire); stipules fully amplexicaul, connate.
Pistillate inflorescences globose-capitate; perianth 2–3-lobed, stigma tongue-shaped.
Trees or shrubs, often with stilt-roots.
Seed large, without endosperm.
Inflorescences bracteate.
