Tree 8-30 m. Twigs subterete or slightly flattened, with two raised lines or shallow wings up to 2 mm high joining the petioles, 4—8(—15) mm diameter, smooth or finely striate, early glabrescent, hairs 0.1 mm or less, stem hollow and in places somewhat swollen and with a longitudinal opening (ant-swellings) or without distinct ant-swellings; older twigs coarsely striate, sometimes transversely cracked; lenticels conspicuous or inconspicuous. Leaves membranous or (thinly) chartaceous, (obovate-)oblong or oblong-lanceolate, 18-52 by 6-22 cm, base rounded or shallowly cordate, apex acute-acuminate; upper surface olivaceous or dull brown, lower surface grey, glabrescent, indumentum minute; not conspicuously papillose; dots absent; midrib faintly raised above, lateral nerves 20-30(-35) per side, at 60-80° to the midrib, ± flat above, lines of inter-arching not very prominent, venation often scalariform, distinct or indistinct; petiole 15-20 by 2.5-5 mm; leaf bud 10-20 by 3-4 mm, hairs appressed, 0.1-0.2 mm. Inflorescences among the leaves or below, of the Knema-type, indumentum minute, glabrescent; in male: brachyblast simple or forked, scar-covered, wart-or worm-like, to 1.5 cm, sessile or with peduncle to 1 cm (var. brevipilis), with 1-5 flowers apically, buds of different sizes; female inflorescences: subsessile, l-3(-5)-flowered; flowers minutely pubescent, ± glabrescent. Male flowers: pedicel 6-15 by 0.8-2 mm, bracteole ovate(-elliptic), 3-5.5 mm, persistent; buds ovoid-lanceolate, 10-13 by 3.5-6 mm, apex narrowed, blunt, base ± rounded, cleft c. 1/8, lobes 1.5 mm long, 0.2-0.3(-0.5) mm thick. Androecium 6.5-9 mm; androphore 1.5-2 by 0.8-1 mm, subglabrous or minutely pubescent in the lower 2/3-3/4; synandrium narrowed to the apex, 3.5-6 by 0.8-1.5 mm, thecae 12-14, sterile apex subacute, narrow, 0.5-1 mm. Female flowers: pedicel 3-5 by 2-2.5 mm, bracteole broadly rounded, 2 mm; buds ± leathery, ovoid-oblong, ± narrowed to the apex, (5-)7-9 by 3-4.5 mm, lobes 1.5-2 mm; ovary ovoid-oblong, finely pubescent, including stigma, 5-6 mm. Fruits 1-6 per infructescence, (sub)sessile, ± fusiform or ellipsoid, 5-7 by 3.5-4.5 cm, minutely mealy brown pubescent, glabrescent; pericarp 10(—15) mm thick, coarsely wrinkled; seeds ellipsoid, (2.5-)3-4 cm; fruiting pedicel (4-)5-12 by 3-5 mm, with some lenticels.
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