Tree 6-36 m. Twigs 2-5(-10) mm diameter, usually subangular with two raised lines joining the petioles, but not winged, frequently ± hollow, but without ant-swellings; (early) glabrescent, hairs palish, less than 0.1 mm; older twigs with small, ± concolorous lenticels. Leaves chartaceous, elliptic-oblong or oblong-lanceolate, 15-35(-40) by 4-12 (-15) cm, base cuneate, rounded, or shallowly cordate, apex acute-acuminate; upper surface olivaceous, lower surface olivaceous or pale brownish, early or late glabrescent, hairs weak, dense or scattered, stellate, 0.1 mm or less; not or ± papillose; dots absent; midrib flat above, lateral nerves 15-25 per side, flat, at 45-60° to the midrib, lines of interarching and venation indistinct; petiole 15-25 by 2-5 mm; leaf bud 10-15 by 2-4 mm, hairs 0. l(-0.2) mm. Inflorescences between the lower leaves or below, as in Knema; in male: a simple or up to 4-branched, to 1 cm (rarely to 3 cm) long, scar-covered, glabrescent brachyblast, rarely to 5 mm pedunculate, with subumbel(s) of 5-10 flowers, buds of various sizes; in female similar, short, (sub)sessile, with a 5-10-flowered cluster; flowers with mealy hairs 0.1 (-0.2) mm. Male flowers: pedicel 4-6 by 1 mm, bracteole rounded, 2(-3) mm, (sub)persistent; buds thinly leathery, (subglobose), broadly ellipsoid, or ovoid, 4-6.5 by 4-5 mm, cleft l/4(-l/3), lobes 1.5-2 mm long, (0.3-)0.5 mm thick. Androecium 3-4(-5) mm; androphore (0.5-)l(-1.5) by 0.5-0.6 mm, hairs minute, in lower 1/4 or lower; synandrium ovoid-oblong, 2-4 by 1 mm, thecae c. 16, the sterile apex ± acute, 0.1-0.3 mm. Female flowers: pedicel 2-3 (-4) by 2 mm, brac-teole 2 mm; buds ovoid, 5-6 by 4-5 mm, lobes 1 mm long, ovary ovoid-oblong, 4 by 3 mm, minutely pale brown pubescent, stigma 2 (^4)-lobed, 1 mm. Fruits subsessile, l-4(-5) per infructescence, (sub-globose-) ellipsoid, 2.5-4.5(-5, see note 2) by 2-3 cm, the apex often shortly pointed, hairs thinly set, mealy, brown, 0.1 (-0.3) mm, ± gla-brescent, with some indumentum remaining near the base and in depressions; pericarp 3-5(-7) mm thick; seeds (broadly) ellipsoid, 2-3 cm; fruiting pedicel 3-7(-10) mm, smooth or with few lenticels, bracte-ole scar subapical.
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A large tree up to 40 m tall with a straight trunk and buttress roots. Twigs have 2 raised lines along them between the leaves. The leaf stalk is 3 cm long and 0.3 mm wide. The leaf is 20-35 cm long x 5-13 cm wide. The base of the leaf is rounded and the tip tapers. The midrib and veins are sunken on the top of the leaf and raised underneath. The veins are almost parallel with fine networks between them. The male flower cluster is a woody scar covered knob. The male flowers have stalks 6 mm long. The female flowers are similar to the male flowers but with thicker shorter stalks. The fruit are oval and 3-6 cm long by 2-4 cm wide. Fruits are yellow with a red covered seed inside. The seed is 3 cm long by 1.5 cm wide.
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