Tree 5-40 m. Twigs 1.5-3 mm diameter, early glabrescent, hairs greyish, less than 0.1 mm (in NE Borneo to 0.5 mm); older twigs (blackish) brown, ± flaking, sporadically with a few lenticels. Leaves membranous or chartaceous, rarely (at c. 1000 m) sub-coriaceous, elliptic-oblong to oblong-lanceolate, 7-24 by 1.5—8(—10) cm, base rounded or attenuate, apex acute-acuminate; upper surface olivaceous(-brown), lower surface early glabrescent, hairs scale-like, less than 0.1 mm; not papillose; dots absent; midrib slightly raised above, lateral nerves 11-16 per side, at (30-)45(-80)° to the midrib, above flat to sunken, below rather flat or raised, lines of interarching and venation indistinct; petiole 11-25 by (0.5-)l-2 mm; leaf bud 7-12 by 1-2 mm, hairs 0.2(-0.5) mm. Inflorescences between the leaves or below, paniculate, pedunculate, glabrescent; in male: 2-8 cm long, peduncle slender or stout, 2-30(-40, see note 1) mm long, branches to 20 mm, central axis absent or up to 40 mm, with 1 or 2 subumbels of 5-15 flowers, buds of various sizes; bracts 3-4 mm, caducous; female inflorescences: reduced, sometimes ± sessile, 1-6-flowered; flowers glabrescent, hairs ± woolly, 0.1-0.3 mm. Male flowers: pedicel slender, 4-12 mm, bracteole 1.5-3 mm, subpersistent or caducous; buds ovoid (-ellipsoid), 4-6(-7) by 2-4(-6) mm, cleft (1/4-) 1/3-1/2, lobes membranous or perga-mentaceous, 0.2-0.3 mm thick. Androecium 3(-6) mm; androphore cylindrical or somewhat swollen, about as broad as the synandrium, 1.2-2(-3) mm long, with hairs 0.1-0.2 mm in the lower half; synandrium ellipsoid-lanceolate, 2-3 by 0.6-1 mm, thecae 10-16, ± contiguous, sterile apex 0.1-0.3 mm, entire or shallowly lobed, glabrous. Female flowers: pedicel 1-2 or 3-7 mm long (see note 1); buds ovoid, 4-4.5 by 3.5-4 mm (in galled flowers larger); ovary long-ovoid, 3 by (1-)1.5 mm, hairs dense 0.1 mm or less. Fruits usually single, ellipsoid or ovoid-ellipsoid, 3.5-6(-8) by 2.5-3.5(-4.5) cm, glabrescent, hairs dense, mealy, grey-brown, 0.2 mm; pericarp (4-)5-10 mm thick; seeds ellipsoid, 2-4(-5) by l-2(-2.3) cm; fruiting pedicel either short and thick, 1-3 mm long (Java), or slender, 5-15 mm long (Borneo) (see note 1).
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