Tree 5-20 m. Twigs subterete or ± flattened, 4-10 mm diameter, with wings 1-2 mm high, early glabrescent, hairs 0.1 mm, hollow, and usually with ant-swellings with longitudinal slit-like opening; older twigs with scattered lenticels. Leaves (sub)chartaceous or subcoriaceous, (ob)ovate-oblong or oblong(-lanceolate), 15-40 by 5-14 cm, base cuneate, rounded, or subcordate, apex acute-acuminate; upper surface olivaceous or brown, lower surface grey(-brown), glabrescent, hairs scale-like, scattered, less than 0.1 mm, or indumentum dense, short (subsp. velata); conspicuously irregularly papillose; dots absent; midrib faintly raised above, lateral nerves 20-25 per side, at 50-70° to the midrib, flat or sunken, lines of interarching and venation not prominent; petiole 15 by 3-4 mm; leaf bud 10-15 by 2-3 mm, hairs 0.1 mm. Inflorescences of the Knema-type, (sub)sessile scar-covered brachyblasts, subglabrous, to 10 mm long, simple or 2-branched; in male: with subumbels of (5-) 10-15 flowers, buds unequal in size; in female: 5-8-flowered, sometimes bearing some fruits of a previous season; flowers with hairs 0.1 mm or less. Male flowers: pedicel (7-) 10-12 by (0.6-)l mm, bracteole ± 3-topped, 2-3 by 2-3 mm, persistent; buds ovoid-oblong, 6.5-7.5 by 3.5-4 mm, apex blunt and faintly triangular or not, base (broadly) rounded, cleft 1/4-1/3, lobes 1.5—2.5(—3) mm long, 0.2-0.3 mm thick. Androecium 5.5-6.5 mm; androphore (1—)1.5—2 by 0.5-0.8 mm, hairs 0.1 or less ± in the lower half, sometimes only at the base; synandrium 4-5 by 0.8-1 mm, rather acute, thecae 12-14, sterile apex 0.2-0.3 mm. Female flowers: pedicel 3-5 mm, bracteole 2 mm, buds ± ovoid, 5 by 3.5 mm, lobes 1.5 mm; ovary ± ovoid, 3 mm, finely pubescent. Fruits solitary or 2-4 per infructescence; (broadly) ovoid(-ellipsoid), 2.5-3.5 by (1.5-)2(-2.5) cm, apex ± rounded and short-apiculate, hairs rust-coloured, 0.1 (-0.2) mm; pericarp 2 mm thick; seeds ellipsoid, 2 cm; fruiting pedicel ± stout, short or long, 3-10 by 3-4 mm, with a few lenticels.
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