Tree 3-10(-25) m. Twigs ± flattened, with two raised lines or wings to l(-2) mm high between the petioles, usually hollow and with ant-swellings (these faint or absent in var. hagensis), (3-)4-8(-10) mm diameter, glabrescent, hairs 0.1 mm or less; older twigs sometimes shallowly fissured, lenticellate. Leaves membranous, chartaceous or (sub)-coriaceous, elliptic(-oblong), 15-40 by 5—13(—16) cm, base cuneate or (narrowly) rounded, sometimes shallowly cordate, apex acute-acuminate; upper surface olivaceous or brown, lower surface grey-brown, glabrous or pubescent; neatly or irregularly papillose or not papillose; dots absent; midrib flat above, lateral nerves (13-) 17-25 per side, at 50-80° to the midrib, flat or sunken, lines of interarching and venation distinct or not; petiole (10-)15-25 by (2-)3-5 mm; leaf bud 10-15(-20) by 2-3 mm, hairs 0.1(-0.2) mm. Inflorescences between the leaves or rarely below, of the Knema-type: a sessile or to 4 mm pedunculate (or 5-20 mm in var. pedunculata), woody, wart-or worm-like, scar-covered brachyblast, 10(-20) by 5 mm, sometimes 2-or 3-armed, pubescent or glabrescent, ending in few-or many-flowered subumbels (in male to 20-flowered, in female 1-3-flowered), the buds either ± equal in size or of various sizes; flowers with hairs 0.1-0.2 mm, sometimes (sub)glabrous. Male flowers: pedicel 8-16 by (0.6-)l mm, bracteole sometimes ± 3-topped, 2(-3) mm, persistent or caducous, (sub)apical; buds (mostly) membranous, (ellipsoid-)oblong or ± tubiform, 8-15 by (2-)2.5-5 mm, apex ± obtuse or subacute, not or faintly angled, cleft 1/6 or less, lobes 1.5-2 mm long, 0.2-0.5 mm thick. Androecium slender or ± stout, 7-12 mm; androphore 2-6 by 0.4-1 mm, subglabrous or variable with scattered hairs less than 0.1 mm, either only at the base or in the lower half; synandrium (4-)5-7 by 0.5-1.5 mm, tapering into acute sterile apex 0.5-1 mm long, thecae 8-24. Female flowers: pedicel 1—5(—7) mm, bracteole 2-3 mm, (sub)apical; buds ovoid-oblong, narrowed to the apex, 4-7(-8) by 2.5-3.5 mm, lobes 1-1.5 mm; ovary narrowed to the apex, 3-3.5 mm long, minutely pubescent, stigma 1.5(-2) mm. Fruits solitary or in clusters of up to 6, subglobose, ellipsoid or broadly ovoid, 1.5-2 (-2.5) by 1-1.5 cm, narrowed to the apex, with the style-remnant remaining as a short apiculum, (fruits 1-3 per cluster, 2.5-3 cm long, with pedicel to 10 mm long in var. paucifructa), hairs persistent, mealy, rusty, 0.1-0.3(-0.4) mm, rarely glabrescent, sometimes lenticels in the basal part; pericarp 2(-3) mm thick; seeds (broadly) ovoid or ellipsoid, 1.5(-2) cm; fruiting pedicel 2-5(-10) by l-2(-2.5) mm, generally with conspicuous raised lenticels.
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