Tree 3-20(-25) m. Twigs usually markedly (2-or) 3-angular, 1—2.5(—3) mm diameter, early glabrescent, hairs 0.1 (-0.2) mm or less; older twigs with inconspicuous lenticels. Leaves membranous or thinly coriaceous, elliptic(-oblong) or oblong-lanceolate, base attenuate, apex acute(-acuminate), 6-16 by (1.5—)2.5—5.5 cm; upper surface olivaceous or (bright) brown, lower surface slightly paler, early glabrescent, hairs weak, scattered, 0.1 mm or less; not papillose; dots absent (minute dots sometimes present in young leaves, see note 1 under subsp. warburgii)', midrib raised above, lateral nerves 10-20 per side, at (45-)60-70(-80)° to the midrib, flat or sunken and inconspicuous above, little raised below, lines of interarching and venation distinct or faint; petiole 7-15 by 1-2 mm; leaf bud 7-10 by 1.5-2 mm, hairs (0.2-)0.3-0.4 mm. Inflorescences between the leaves or below, of the Knema-type: a sessile or to l(-2) mm pedunculate simple or forked scar-covered brachyblast to 4 mm long, late glabrescent (hairs 0.3 mm), in male: subumbel of 3-6 flowers, buds of various sizes; bracts minute; in female: with 1-4 flowers, almost equal in size; flowers often partially glabrescent or subglabrous, hairs 0.1 mm or less. Male flowers: pedicel slender, 5-10 by 0.5-1 mm, bracteole ± ovate, 1-1.5 mm, (persistent or) caducous, apical or to 2.5 mm below apex; buds oblong-lanceolate, ± fusiform (with base attenuate) or tubiform, the upper 1/3 narrowed, acute, ± angled, 8-14 by 1.5-3 mm, cleft c. 1/8, lobes 1-2 mm long, 0.2-0.3 mm thick. Androecium slender, 5.5-9 mm long; androphore (2-)2.5-4.5 by 0.5-0.7 mm, hairs weak, glossy, 0.2-0.3 mm long; synandrium 3.5-5 by 0.7-0.8(-l) mm, thecae 12-16, sterile apex ± acute or blunt, 0.1-0.5 mm long, or absent. Female flowers: pedicel 9-15 by 0.8-1.5 mm, bracteole scar 3-6 mm below apex; buds ovoid-oblong, narrowed in the upper part, 7.5-9 by 3.5-4 mm, lobes 1 mm; ovary ovoid-oblong, 6 by 2-3 mm, hairs 0.2-0.3 mm. Fruits solitary or 2 or 3 together, long-stiped, often axillary to leaves, broadly ellipsoid, sometimes fusiform, (1.5-)2-4 by 1.5-2.5 cm, apex rounded or narrowed, often somewhat acute with up to 2 mm long beak, base tapered or ± rounded and contracted into 2-5(-7) mm long pseudostalk, hairs dense, mealy, brown, 0.1-0.2 mm or less; pericarp 1.5-3 mm thick; seeds ellipsoid, (1.5—)2 cm, attached (up to c. 5 mm) above the base of the fruit rendering seeds and aril lobes generally in a transverse, not erect, position (see note 2 under subsp. warburgii); fruiting pedicel slender, 10-20(-25) by 1.5-3 mm, sometimes with few lenticels, bracteole scar at about 1/4 to halfway from apex.