Plants medium-sized to large, shelf-forming, light to dark green or sometimes yellowish, usually shiny. Stems creeping, irregularly branched; paraphyllia present or absent. Secondary stem and branch leaves (central) erect, erect-spreading, or spreading, sometimes secund, ovate, obovate, oblong, oblong-ligulate, or oblong-lanceolate, asymmetric, flat to strongly undulate; margins entire to serrate, teeth recurved; apex obtuse to acuminate; costa double and short, sometimes single or ecostate; basal laminal cells linear to rectangular, walls pitted or not. Sexual condition autoicous or dioicous; perichaetial inner leaves oblong, oblong-lanceolate, or ovate-subulate. Seta (0.1-)0.2-1.2 cm. Capsule cylindric, oblong, or globose; exostome teeth lanceolate, smooth to densely papillose, somewhat cross striate basally; endostome basal membrane low to high, segments smooth to densely papillose. Spores 12-39 µm.
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