Herbs annual, biennial, or perennial with a woody caudex. Trichomes stalked or subsessile, 1-or 2-forked sometimes also simple. Stems erect or prostrate, usually several from base. Basal leaves petiolate or sessile, rosulate or not, entire, dentate, or pinnatisect. Cauline leaves often similar to basal ones, rarely absent. Racemes few to several flowered, ebracteate or rarely bracteate. Fruiting pedicels erect, ascending, or divaricate. Sepals ovate or oblong, erect or spreading, rarely reflexed, glabrous or pubescent, base of lateral pair not saccate. Petals white or pink, sometimes yellowish, longer or rarely shorter than sepals; blade obovate, spatulate, or oblanceolate, apex obtuse or retuse; claw distinct or not. Stamens 6, tetradynamous; filaments rarely dilated at base; anthers ovate or narrowly oblong, apex apiculate or obtuse. Median glands absent; lateral ones semilunar or 1 on each side of lateral stamen. Ovules (8-)16-36(-44) per ovary. Fruit dehiscent siliques, linear or rarely linear-oblong, terete or 4-angled, sessile; valves with branched trichomes sometimes mixed with simple ones, rarely glabrescent, torulose; replum rounded; septum complete; style obsolete or to 1 mm; stigma capitate, entire or 2-lobed. Seeds uniseriate, wingless, oblong or ovate, plump; seed coat reticulate, not mucilaginous when wetted; cotyledons incumbent.