Habit, rhizome morphology. Runners 1-2 mm thick, branching angle divaricate. Scales on runners dense, spreading or squarrose. Tubers absent. Fronds to 200 cm long (or much longer), 15-19 cm wide, stipe 30-35 cm long. Lamina base strongly reduced, tapering over 20-60 cm, basal pinnae 1-1.2 cm long, 2.5-6 cm distant, middle pinnae slightly to strongly falcate (mostly the cauda only is falcate). Sterile pinnae 3-13 by 0.8-2.8 cm (usually distinctly triangular, narrowed gradually from base to apex), leathery, base strongly unequal, basiscopic base cordate (strongly, sometimes somewhat auriculate), acroscopic base cuneate, truncate, rounded or cordate (always less distinctly cordate than the basiscopic base), not or slightly auricled (usually distinctly dilated, but not auricled), margin in basal part finely crenate, towards apex crenate, apex rounded, obtuse or acute. Fertile pinnae 3.2-14 by 0.7-1.8 cm, differing from sterile pinnae in the margin being incised between sori but not beyond the attachment. Indument. Basal scales pseudopeltate, spreading, 5.5 by 1 mm, central part rufous, shining (translucent), hyaline margin absent, marginal glands absent, margin in basal part fimbriate, acumen fimbriate, transition to rhachis scales abrupt or basal scales persisting to high up. Rhachis scales sparse or dense, with a well-developed protracted ciliate acumen, appressed or spreading, light brown or rufous. Scales on lamina absent. Hairs on lamina absent or present, short, dense. Sori marginal, 14-34 pairs on fully fertile pinnae, elongated, not impressed. Indusium broad, attached at broad base.