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Fls perfect; tep 2 + 2, the inner broader; stamens 6, about equaling the tep; ovary flattened, with 2 short, divergent styles, the stigmas tufted; fr a flat achene surrounded by a broad, radially striate wing, cordate at base, subtended by the persistent, slightly enlarged tep, the 2 outer spreading, linear-oblong, the inner appressed, obovate-spatulate; perennial glabrous herbs with a stout, multicipital root, bearing several basal and few cauline lvs on long petioles, the blade broadly cordate to reniform; fls minute, in branching panicles; foliage acid. Monospecific.
Herbs perennial or weakly defined subshrubs, sometimes dioecious. Rhizomes large. Stems erect, sparsely to densely branched. Leaves simple, alternate, petiolate; leaf blade reniform, orbicular-reniform, or orbicular-cordate, margin entire or nearly so; ocrea tubular, membranous, margin entire, apex oblique or truncate. Inflorescence terminal, paniculate. Pedicel articulate. Flowers bisexual or unisexual. Perianth persistent, accrescent in fruit. Stamens 6. Ovary compressed; styles 2; stigmas penicillate. Achenes biconvex, ovoid, margin broadly winged.
