Plants coarse. Stems with paraphyllia few to many, spread on stem or often in ± transverse bands. Stem leaves gradually narrowed to apex, concave; alar cells many, walls thin or slightly incrassate, region not expanded marginally, reaching from margin to costa or almost so; rarely some single basal medial laminal cells abaxially prorate, rarely single cells with papillae. Branch leaves smaller. Perichaetia with inner leaves gradually narrowed to apex, plicate, margins denticulate distally [sometimes with scattered cilia], apex acuminate, laminal cells smooth.
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