Plants medium-sized to large, green, yellow-green, or brownish yellow. Stems sparsely pinnate or irregularly branched; hyalodermis absent, central strand weak or absent; paraphyllia present, ; rhizoids or rhizoid initials on stem or abaxial costa insertion, often forming tomentum, strongly branched, at least partly warty-papillose; axillary hair distal cells 2-6, hyaline. Stem leaves not recurved or squarrose, strongly falcate to ± straight, ovate or triangular-ovate, plicate or strongly so, longer than 1 mm; base ± decurrent; margins plane or sometimes reflexed basally, distinctly denticulate usually only proximally, limbidia absent; apex long or more rarely short, ; costa strong, single, ending in apex, sometimes weaker and shorter; alar cells differentiated, short-to long-rectangular, inflated, hyaline or yellow when young, region usually well differentiated, transversely triangular or narrowly so; medial laminal cells linear, ; marginal cells 1-stratose. Sexual condition dioicous. Capsule horizontal, cylindric, curved; peristome perfect; exostome margins dentate distally; endostome cilia 2-3(-4), well developed, nodose. Spores 10-23 µm.
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