Paspalum distichum L.

Water couch (en), Paspale à deux épis (fr), Paspale distique (fr)

Species

Angiosperms > Poales > Poaceae > Paspalum

Characteristics

Perennials, with numerous long-creeping branching stolons to 3 mm diam., rooting at nodes. Leaf-sheath submembranous, glabrous. Ligule 1-1.6 mm, erose, truncate. Collar hairs few, long, tubercle-based. Leaf-blade 2.5-9 cm × 2-8 mm, rather soft, flat, linear-lanceolate, curved at base, abaxially glabrous with obvious midrib, adaxially with scattered, long, fine tubercle-based hairs; margins scabrid, gradually narrowed to acuminate scabrid tip. Culm 7-35-(45) cm, erect or geniculate-ascending, compressed, nodes long-hairy, internodes glabrous. Panicle of (1)-2-(3) erect to spreading racemes, ± digitate at culm apex. Racemes 2-5 cm, sometimes one or both shortly pedunculate; axils with a few long white hairs; rachis 1-2 mm wide, concave, narrowly winged with scabrid margins, bearing 2 rows of single or paired, subsessile or shortly pedicelled spikelets; pedicels scabrid; paired spikelets usually near centre of rachis. Spikelets c. 2.5-3.5 mm, imbricate, elliptic, acute, light green. Lower glume a minute scale rarely to 1 mm, or 0, upper = spikelet, 5-nerved, minutely appressed-puberulent. Lower floret: lemma ≈ upper glume and slightly less membranous, 3-5-nerved, glabrous; palea rarely present, narrow, short, hyaline, 2-keeled. Upper floret: lemma 2-3 mm, faintly 5-nerved, cartilaginous-indurate, finely punctulate-striolate, broad-elliptic, acute, light cream, apex often with tuft of short hairs; palea ≈ lemma, hyaline margins only slightly widened near centre; anthers 1-1.7 mm, dark purple; stigmas dark purple; caryopsis 1.5-2 mm.
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Perennial, 100-300 mm high; hydro-to hygrophyte; mat-forming as rhizomatous or stoloniferous and rooting at nodes; culm nodes pubescent. Leaf blade 20-220 x 3-8 mm; ligule an unfringed membrane. Inflorescence digitate to subdigitate, usually 2, secund racemes; rachis dorsally flat, with a raised midrib on front, bearing single spikelets alternately on either side of midrib in 2 or 4 rows; lower glume abaxial, upper lemma facing rachis. Spikelets 2.5-3.5 x 1.3-1.5 mm, lanceolate, dorsiventrally compressed, apex acute, glabrous, awnless, green; glumes unequal; lower glume usually a small triangular scale, or up to 2/3 as long as spikelet to absent (all 3 states can occur on same inflorescence); upper glume minutely hairy especially towards apex, 3-nerved. Florets 2; lower floret sterile, lemma equal, similar to upper glume but flat on back, a prominent median nerve present; upper floret bisexual, lemma hard, obtuse, entire, median nerve present, glabrous, smooth, pallid at maturity, margins narrow and inrolled, clasping only edges of palea; anther 1.0-1.6 mm long. Flowering time Nov.-May.
Stoloniferous; culms 2–5 dm, ascending from a creeping base with conspicuous sheaths; lf-blades relatively short, 2–8 cm × 2–5 mm, ciliate at base; racemes 2, one sessile, the other short-stalked, ascending and often incurved, 2–5 cm, the rachis 1–1.5 mm wide; spikelets solitary, 2.6–3 mm, half as wide, plump, planoconvex, ovate; first glume often developed, triangular, to 1 mm; second glume 5-veined, minutely pubescent, this and the sterile lemma somewhat leathery; 2n=40, 48, 60. Swamps, wet ground, and waste places, often in coastal salt-marshes; mainly in subtropical and warm-temperate regions throughout the world, n. in our range to Ky. and se. Va., and adventive to N.Y. There is some controversy as to whether the name P. distichum should apply to our sp. or to the closely related, more tropical plant usually called P. vaginatum Sw.; in the latter case, our plants take the name P. paspalodes (Michx.) Scribn.
Perennial with rhizomes and stolons. Culms 20–50 cm tall, nodes usually pubescent. Leaf sheaths keeled, glabrous, margins ciliate; leaf blades linear, 5–10 × 0.3–0.7 cm, glabrous, apex acute; ligule 2–3 mm. Inflorescence of 2(–3) racemes arising together or separated by a short axis; racemes 3–7 cm; spikelets single, in 2 rows; rachis straplike, 1.5–2 mm wide. Spikelets pallid, obovate-oblong, plano-convex, 3–3.5 mm, acute; lower glume vestigial or a narrow triangular scale up to 1/2 spikelet length or more; upper glume papery, 3–5-veined with distinct middle vein, loosely appressed pubescent; lower lemma 3–5-veined, usually glabrous; upper lemma pale green, almost equal to spikelet, cartilaginous, apex apiculate and minutely pubescent. Fl. and fr. May–Sep. 2n = 40, 48, 60.
Perennial 100-300 mm high, hydrophyte; mat-forming as rhizomatous or stoloniferous and rooting at nodes; culm nodes pubescent. Leaf blade 20-220 x 3-8 mm; ligule 1.5-3.5 mm long, conspicuous. Inflorescence usually of two racemes; spikelets arranged in two rows, secund. Spikelet 2.5-3.5 x 1.3-1.5 mm, lanceolate, green, apex acute; lower glume usually a small triangular scale up to 2/3 as long as spikelet to absent (all 3 states can occur on the same inflorescence); upper glume 3-nerved, minutely hairy especially towards apex; lower lemma with a prominent midrib; upper lemma smooth, pallid at maturity; anther 1.0-1.6 mm long.
Perennial; up to 0.3 m high; hydrophyte; rooting at nodes; rhizomatous or stoloniferous (matforming). Leaf blades 20-220 x 3-8 mm. Flowers: inflorescence digitate; racemes 2; spikelets 2.5-3.5 x 1.3-1.5 mm; arranged in 2 rows; glabrous; narrowly obovate; apices acute; lower glume usually developed into a small triangular scale or sometimes absent; upper glume minutely hairy.
Creeping, hydrophytic perennial, up to 300 mm tall. Leaves linear; ligule an unfringed membrane. Inflorescence digitate with 2 or 3, 1-sided racemes. Spikelets in 2 rows, 2.5-3.5 mm long, green, ovate, minutely hairy all over, hairs short, awnless; 2-flowered; upper glume as long as to longer than and covering spikelet; lower floret sterile, soft; upper lemma hard, pallid.
Perennial, hydrophyte (rooting at nodes), or rhizomatous or stoloniferous (mat-forming), up to 300 mm tall. Leaf blades 20-220 mm long, 3-8 mm wide. Spikelets 2.5-3.5 mm long, 1.3-1.5 mm wide. Racemes usually 2; spikelets arranged in 2 rows, glabrous, lanceolate, tips acute; lower glume usually developed into a small triangular scale or sometimes absent.
Perennial hydrophyte to 30 cm. Leaves linear. Spikelets in 2 or 3, digitate, spike-like branches, secund in 2 rows, 2.5-3.5 mm long, upper glume minutely hairy.
Inferior glume present on at least some spikelets as a tiny scale. Superior glume somewhat coriaceous, nearly always obscurely pubescent.
Racemes paired (rarely 3), each 2–8 cm. long, the spikelets borne singly in 2 rows on a rhachis 1–2 mm. wide.
A grass. It keeps growing from year to year. It has underground stems or rhizomes and also runners.
Inferior lemma somewhat coriaceous. Superior lemma pallid at maturity.
Spikelets 3–4 mm. long, ovate, plump, green.
Creeping stoloniferous perennial.
Culms 8–60 cm. high.
Life form perennial
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality hermaphrodite
Pollination anemogamy
Spread epizoochory
Mature width (meter) -
Mature height (meter) 0.2 - 0.3
Root system rhizome
Rooting depth (meter) 0.8
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c4

Environment

Sands and muds near the seashore; saline soil; swamps; wet marshlands; polluted shallow water; along irrigation ditches, etc at elevations up to 1,700 metres.
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It is a tropical plant.
Light 7-8
Soil humidity 8-11
Soil texture 1-6
Soil acidity 2-9
Soil nutriment -
Hardiness (USDA) 8-10

Usage

Uses animal food environmental use fodder food gene source medicinal poison vertebrate poison
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by seedlings.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -12
Optimum temperature (C°) 20 - 30
Size -
Vigor -
Productivity -

Images

Habit

Paspalum distichum habit picture by Manuel Barón (cc-by-sa)
Paspalum distichum habit picture by Renaud Brochiero (cc-by-sa)

Leaf

Paspalum distichum leaf picture by Sonia Henno (cc-by-sa)
Paspalum distichum leaf picture by Kay (cc-by-sa)
Paspalum distichum leaf picture by histoffe (cc-by-sa)

Flower

Paspalum distichum flower picture by Mehmudul Mehmudul Islam Ansary (cc-by-sa)
Paspalum distichum flower picture by Mehmudul Mehmudul Islam Ansary (cc-by-sa)
Paspalum distichum flower picture by Manuel Barón (cc-by-sa)

Fruit

Paspalum distichum fruit picture by Robin Boyd (cc-by-sa)

Distribution

Paspalum distichum world distribution map, present in Åland Islands, Albania, Andorra, Austria, Bangladesh, Bulgaria, Bahamas, Bermuda, Bolivia (Plurinational State of), Brazil, Canada, China, Cook Islands, Comoros, Costa Rica, Cuba, Cyprus, Dominica, Algeria, Ecuador, Egypt, Spain, Fiji, France, Georgia, Greece, Guatemala, Guyana, Honduras, Haiti, Indonesia, Iran (Islamic Republic of), Iraq, Italy, Jamaica, Japan, Libya, Lesotho, Morocco, Moldova (Republic of), Maldives, Myanmar, Mauritania, Mauritius, Namibia, Norfolk Island, Nicaragua, Nepal, New Zealand, Oman, Pakistan, Peru, Philippines, Palau, Puerto Rico, Korea (Democratic People's Republic of), Portugal, Paraguay, Réunion, Saudi Arabia, Singapore, Suriname, eSwatini, Turks and Caicos Islands, Togo, Tunisia, Tuvalu, Taiwan, Province of China, Uruguay, United States of America, Venezuela (Bolivarian Republic of), Viet Nam, Samoa, Yemen, South Africa, and Zimbabwe

Conservation status

Paspalum distichum threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:184275-2
WFO ID wfo-0000887584
COL ID 4DW5G
BDTFX ID 48018
INPN ID 112483
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Anastrophus paspalodes Digitaria paspalodes Milium distichum Panicum digitaria Paspalum bracteatum Paspalum paspalodes Paspalum paucispicatum Digitaria disticha Panicum paspaliforme Paspalum elliottii Dimorphostachys oajacensis Panicum fernandezianum Paspalum berteroanum Paspalum glaucifolium Paspalum paspaliforme Paspalum polyrrhizum Paspalum schaffneri Panicum polyrrhizum Paspalum chepica Paspalum digitaria Paspalum michauxianum Paspalum fernandezianum Milium paspalodes Paspalum distichum subsp. paspalodes Paspalum distichum var. digitaria Paspalum distichum var. indutum Paspalum distichum var. longerepens Paspalum distichum var. microstachyum Paspalum distichum var. paspalodes Paspalum vaginatum var. pubescens Paspalum paspalodes var. paspalodes Paspalum distichum subsp. paucispicatum Paspalum oajacense Paspalum oaxacense Paspalum distichum