Annuals or, rarely, biennials or perennials; not scapose; pubescent or glabrous, trichomes short-stalked, forked or subdendritic, often mixed with coarse, simple ones. Stems erect, ascending, decumbent, or procumbent, unbranched or branched. Leaves basal and cauline; petiolate or sessile; basal rosulate, petiolate, blade margins entire, dentate, or pinnately lobed; cauline sessile, blade (base usually auriculate or sagittate, sometimes amplexicaul), margins dentate to lobed. Racemes (several-flowered), often strongly elongated in fruit, (rachis straight). Fruiting pedicels ascending to divaricate or recurved, (straight or slightly curved), slender. Flowers: sepals (deciduous), suberect, ascending, or spreading, oblong, (equal), lateral pair not saccate basally, (margins membranous); petals yellow or white, broadly obovate, claw slightly differentiated from blade, (apex rounded, truncate, retuse, or emarginate); stamens (erect), tetradynamous; filaments dilated or not basally; anthers oblong or sagittate, (not apiculate); nectar glands confluent, subtending bases of median stamens, surrounding lateral stamens. Fruits silicles, subsessile or sessile, globose, subglobose, subpyriform, suborbicular, orbicular, elliptic, cordate, or obovoid, terete, flattened, latiseptate, or angustiseptate; valves (papery or leathery), not veined, glabrous or pubescent, (trichomes branched and/or simple); replum not flattened; septum complete, perforated, or reduced to a rim, (membranous); ovules 4-40 per ovary; style distinct, (persistent, slender); stigma capitate, entire. Seeds sub-biseriate, flattened, margined, orbicular, suborbicular, or oval; cotyledons accumbent. x = 7, 8, 9.
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