Trees, dioecious or monoecious. Leaves alternate, distichous on the lateral branches; stipules fully amplexicaul, free; blade entire, pinnately veined. Inflorescences on spurs in the leaf axils, unisexual, discoid-capitate, involucrate. Staminate inflorescences several together, pedunculate, open before anthesis; flowers numerous; tepals 2-4, free or basally connate; stamens 2-4, pistillode absent. Pistillate inflorescences solitary or accompanied by staminate ones, subsessile or pedunculate; flowers several to numerous, free or basally connate; perianth 2-4-lobed to 2-4-fid; ovary free or partly adnate to the perianth; stigmas 2, tongue-shaped or filiform. Fruiting perianth enlarged, fleshy, red(dish); fruit free or partly adnate to the perianth; seed large, without endosperm, cotyledons thick and equal.