Shrubs. Leaves coriaceous, variable. Inflorescence terminal or axillary, cone-like, globular, ovoid or cylindrical; axis woody, sessile or pedunculate; involucral scales usually present, often deciduous; 'cone' scales broad, becoming hardened, persistent; floral bracts absent. Flowers sessile. Tepals separating from base, falling united or separated. Stamens 4, sessile; connective short. Hypogynous glands absent. Ovary sessile; ovules 1 (rarely 2), pendulous; style filiform, straight; pollen presenter fusiform or distinctly enlarged, basally truncated, terminally brush-like, glabrous or hairy; stigmatic surface glabrous, minute. Fruiting 'cones' persistent, sessile or pedunculate. Fruit a small nut, often compressed, ±ovoid to obovoid; style base ±persistent, usually marginally or basally long-haired, sometimes winged.
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