Base of stipe very dark, castaneous distally, scales (near base only) to 2.5 cm long, 1 mm wide at base, distally tiliform; pinnules of larger supra-basal pinnae to 9 × 2 cm with short winged stalks, lobed 2/3-3/4 towards costae, lobes distinctly crenate, obtuse; sinus-teeth broadly triangular, oblique; veins forming costal areoles and a few additional ones below sinuses, costular areoles present only at bases of pinnule-lobes; sori exindusiate, usually about midway between costules and margin, mostly on acroscopic branches of forked veins (none seen on both branches); spores spinulose.