Lower 30 cm of stipe ± persistently covered with scales, the longest 5 cm long, gradually narrowed from a base 3 mm wide; pinnules of supra-basal pinnae commonly 8 × 1.7 cm, lobed 2/3 towards their costae, lobes well-spaced, obtuse, their margins at most sinuous and bearing ± abundant short hairs; costules 4-5 mm apart; veins usually dark on lower surface when dried, forming one series of areoles between costal areoles and sinuses, costular areoles many; sinus-teeth short, broadly triangular and slightly oblique; sori exindusiate, sometimes on both branches of a forked vein, also on anastomosing veins below sinuses; spores with a continuous wing and some cross-wings.