Lianas; stems with 4 phloem arms in cross section, the extreme center hollow; branchlets acutely tetragonal with detachable ribs, without interpetiolar glandular fields; pseudostipules foliaceous or absent. Leaves biternate or tri-ternate or ternately bipinnate or tripinnate, the terminal pinna often replaced by a trifid tendril. Inflorescence an axillary or terminal raceme. Flowers with the calyx cupular, truncate, usually minutely denticulate; corolla white to pale yellow, narrowly campanulate-infundibuliform, the tube glabrous or variously pubescent in upper part; anthers glabrous, the thecae straight, divaricate; pollen grains 3-colpate, the exine microreticulate; ovary oblong, more or less lepidote or pubescent, the ovules 2-seriate in each locule; disc cupular-pulvinate. Fruit an elongate-linear to elongate-oblong compressed capsule, the valves parallel to the septum, more or less smooth, drying blackish, the median nerve incon-spicuous; seeds thin, bialate, the wings membranaceous, brown, poorly demar-cated from the seed body.