Plants epiphytic, on rock, or rarely terrestrial. Stems often long-creeping, usually branched, 1--2.5 mm diam., not whitish pruinose; scales bicolored with dark central region, round to ovate-lanceolate or subulate, clathrate toward center, glabrous or pubescent with reddish brown hairs, margins transparent, erose-denticulate to fringed-ciliate. Leaves monomorphic, widely spaced, not conspicuously narrowed at tip, to 25 cm. Petiole green to dark brown or black, flattened to terete, often grooved proximally and winged distally. Blade oblong-linear to deltate, simple and entire or deeply pinnatifid, not pectinate, with fewer than 25 pairs of segments, not glaucous, with conspicuous peltate scales on abaxial surface; rachis sparsely to densely scaly abaxially, glabrous to sparsely scaly adaxially; scales ovate-lanceolate to spheric, peltate, clathrate toward center. Segments (when present) linear to oblong; margins entire to crenulate; apex rounded. Venation mostly free or complexly anastomosing, with 1--several included veinlets in fertile areoles. Sori often confined to distal 1/2 of leaf, discrete or confluent, circular to oval when immature, borne at ends of single veins or at junction of several veinlets, in 1 row on either side of midrib; indument of ephemeral, peltate scales covering immature sori. Spores smooth with scattered spheric deposits or slightly papillate or verrucose. x = 34, 35, 37.
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