Polystichum braunii (Spenn.) Fée

Braun's hollyfern (en), Polystic de Braun (fr)

Species

Pteridophytes > Polypodiales > Dryopteridaceae > Dryopteridoideae > Polystichum

Characteristics

Plants summer-green. Rhizome erect or ascending, short, densely covered with scales; scales brown, linear. Fronds 40-70 cm; stipe brown at base, 13-21 cm, 4-5 mm in diam. at base, densely covered with scales; scales brown, linear and lanceolate or larger; large scales dense or somewhat sparse, light brown but below middle often blackish brown, shiny, ovate, ovate-lanceolate, or broadly lanceolate, up to 13 × 6 mm, subentire or slightly toothed, long acuminate or caudate. Lamina bipinnate, oblanceolate, 36-60 × 14-24 cm at middle, gradually contracted toward base, lower part sterile, apex acuminate, fertile; rachis without proliferous bulbils, abaxially with dense light brown linear or lanceolate and larger scales; large scales ovate-lanceolate, up to 4.5 mm wide, subentire, caudate or long acuminate. Pinnae 19-25 pairs, alternate, ascendant, shortly stalked, lanceolate, bases asymmetrical, apices acuminate; middle pinnae 7-15 × 2.3-2.8 cm, 1-pinnate; costa narrowly winged, abaxially with linear light brown scales. Pinnules (2-)6-17 pairs, alternate, sessile, oblong, 0.9-1.7 × 0.5-0.9 cm, bases cuneate, decurrent, acroscopic margins entire, sometimes toothed and even shallowly pinnatifid on large plants, shortly or long aristate, apices acute with acute spine; auricles rounded, small, basiscopic margins aristate; basal acroscopic pinnules largest, incised or pinnatifid; both surfaces densely scaly; microscales brownish, filiform; frond texture thinly papery; venation pinnate, lateral veins 5-7 pairs, dichotomous, distinct. Sori large, (1-)3-6 pairs per pinnule, in 1 row along each side of midrib, close to midrib, terminal or sometimes abaxial on veins; indusia present, entire. 2n = 164.
More
Rhizome erect, lvs 3–9 dm; petiole 13–18 cm, densely scaly; blade lanceolate, 7–17 cm wide, acuminate, reduced below, essentially bipinnate; pinnae 30–40 pairs, acuminate, alternate above, the lower subopposite; pinnules 9–15 pairs on the larger pinnae, ovate-deltoid to ovate-oblong, acute, spinulose-denticulate with incurved teeth, the veins of both surfaces bearing linear scales; sori in one row on each side of the midrib; indusium pale, sublobate; 2n=164. Upland woods and rock ledges in circumneutral soil; circumpolar, s. to Mass., N.Y., Pa., Mich., Wis., and n. Minn. The Amer. plants differ slightly from the Eurasian ones and have been segregated as var. purshii Fernald. A hybrid with no. 1 [Polystichum acrostichoides (Michx.) Schott] is P. ×potteri Barrington.
Stems erect. Leaves monomorphic, arching, 2--10 dm; bulblets absent. Petiole 1/8--1/6 length of leaf, densely scaly; scales light brown, gradually diminishing in size distally. Blade broadly lanceolate, 2-pinnate; base narrowed. Pinnae oblong-lanceolate or falcate, proximal pinnae ± rectangular, not overlapping, in 1 plane, 2--10 cm; base oblique except proximal 3--4 pinnae, where auricles not developed; apex acute, subapical and apical teeth same size; microscales filiform to linear, lacking projections, dense abaxially, sparse adaxially. Pinnules ± stalked, short-falcate to oblique-rhombic, acroscopic auricle well developed on proximal pinnules; margins dentate, with slender bristle tips; apex broadly acute. Indusia ciliate. Spores brown. 2 n = 164.
Life form perennial
Growth form herb
Growth support -
Foliage retention evergreen
Sexuality monoecy
Pollination hydrogamy
Spread anemochory
Mature width (meter) 0.5 - 1.0
Mature height (meter) 0.5 - 1.0
Root system rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway c3

Environment

Light 2-5
Soil humidity 2-7
Soil texture -
Soil acidity 2-8
Soil nutriment -
Hardiness (USDA) 7-10

Usage

Uses environmental use medicinal
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by divisions.
Mode divisions
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Polystichum braunii habit picture by manuseitz (cc-by-sa)

Leaf

Polystichum braunii leaf picture by manuseitz (cc-by-sa)
Polystichum braunii leaf picture by manuseitz (cc-by-sa)

Distribution

Polystichum braunii world distribution map, present in Armenia, Austria, Azerbaijan, Belarus, Canada, Switzerland, China, Czech Republic, Germany, France, Georgia, Croatia, Hungary, Iran (Islamic Republic of), Italy, Japan, Kazakhstan, Liberia, Liechtenstein, Latvia, Montenegro, Norway, Poland, Korea (Democratic People's Republic of), Romania, Russian Federation, Saint Pierre and Miquelon, Serbia, Slovakia, Slovenia, Sweden, Turkey, Ukraine, United States of America, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:17206080-1
WFO ID wfo-0001109264
COL ID 4LMNL
BDTFX ID 51899
INPN ID 115046
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Aspidium pilosum Dryopteris braunii Polystichum shennongense Aspidium braunii Polystichum braunii var. purshii Polystichum braunii var. braunii Polystichum braunii subsp. purshii Polystichum braunii