Polystichum Roth

Hollyfern (en), Polystic (fr)

Genus

Pteridophytes > Polypodiales > Dryopteridaceae > Dryopteridoideae

Characteristics

Plants terrestrial. Stems decumbent to erect, stolons absent. Leaves monomorphic (dimorphic in P . acrostichoides ), evergreen. Petiole 1/9--1 times length of blade, bases swollen or not; vascular bundles more than 3, arranged in an arc, ± round in cross section. Blade linear-lanceolate to broadly lanceolate, 1--3-pinnate, gradually reduced distally to pinnatifid apex, somewhat leathery to leathery. Pinnae not articulate to rachis, segment or pinna margins spinulose-toothed (except P . lemmonii ); proximal pinnae (several pairs) usually gradually reduced, sessile to short-petiolulate, bases usually inequilateral with acroscopic lobe; costae adaxially grooved, grooves continuous from rachis to costae; indument of linear to lanceolate scales on costae and sometimes between veins abaxially (microscales), ± glabrous or similarly scaly adaxially (scales forming loosely tangled network over blade and sori in P . dudleyi ). Veins free, forked, rarely ( P . imbricans ) anastomosing. Sori in 1 row (to several) between midrib and margins, round (confluent, covering abaxial surface in P . acrostichoides ); indusia peltate, persistent or caducous [absent]. Spores yellow or brownish to black, with inflated folds. x = 41.
More
Plants perennial, terrestrial, evergreen or summer-green. Rhizome short, erect or ascending, together with basal stipe often covered with scales; scales linear to ovate, rarely with hairs. Fronds caespitose, monomorphic or rarely nearly dimorphic; stipe stramineous to purplish brown, canaliculate adaxially, scaly; distal stipe scales sparser, similar to but smaller than basal stipe scales. Lamina linear-lanceolate, lanceolate, ovate-lanceolate, lorate, or oblong, pinnate, bipinnatifid, or bipinnate, rarely finely divided and tripinnatifid or tetrapinnatifid; frond texture thinly papery, papery, or leathery, with lanceolate, subulate, linear, or ovate microscales abaxially and sometimes also adaxially, acroscopic base often auriculate, rarely auricles indistinct; rachis sulcate adaxially, with or without proliferous bulbils, rarely proliferous bulbils borne terminally on a prolonged flagelliform rachis; venation pinnate, free or rarely anastomosing to form 1 or 2 rows of areoles. Sori orbicular, terminal on veins of pinnae, sometimes abaxial or nearly terminal on veins, indusiate or rarely exindusiate; indusia orbicular, peltate, membranous, entire, erose, or irregularly toothed.
Petioles stout, with 4 or 5 vascular bundles, much shorter than the blades, beset with persistent, brownish, notably dimorphic scales, some broad and often toothed or cleft, others small, appressed, hair-like; blade scaly especially on the rachis and costae, otherwise glabrous or occasionally glandular, pinnate to subtripinnate, the principal pinnae subsessile on a narrow, very shortly petiolulate base, the uppermost ones reduced and confluent; pinnae or pinnules spinulose-toothed, mostly subfalcate; veins obscure, free, mostly once-forked; sori borne on the veins in one or more rows on each side of the midrib; indusium peltate, rotund, persistent; ferns with stout, scaly, erect or short-creeping rhizomes and clustered, firm, evergreen lvs. 135, cosmop.
Terrestrial or lithophytic ferns. Rhizome short, erect, densely scaly; scales variable, becoming erose. Fronds tufted. Stipe densely scaly at the base. Lamina 1-3-pinnate, rarely simple (non-Australian species), often coriaceous; rachis sometimes bearing proliferous buds near the apex; ridges on the adaxial surface of the primary rachis not open to admit the grooves of the secondary rachises; ultimate segments sharply pointed and also sometimes with marginal teeth; veins free. Sori rounded, dorsal or subterminal on veins, indusiate, rarely exindusiate; indusium peltate, persistent or caducous. Spores monolete, ellipsoidal to globose.
Sori dorsal on veins, indusium peltate (rarely absent); sporangia stalked, annulus incomplete, vertical. Spores bilateral, ± oblong, us. tuberculate. Rhizome short, branched, ascending, paleate; stipites densely tufted, paleate. Lamina anadromic, pinnate to decompound, segs us. aristate. Mainly terrestrial ferns with some 180 spp. of wide distribution.
Terrestrial ferns. Rhizomes erect, very scaly. Fronds 1-3-pinnate, scaly; ultimate segments usually pointed; veins free. Sori round, borne on veins in one row either side of midrib; protected by peltate indusia, or indusia 0. Spores monolete.
Fronds elongate, basal pinnae not much enlarged on basiscopic side; rachis often bearing a bud.
Life form perennial
Growth form herb
Growth support -
Foliage retention evergreen
Sexuality -
Pollination -
Spread -
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Mature height (meter) -
Root system rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 7-11

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
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Germination treatment -
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Optimum temperature (C°) -
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Images

Polystichum unspecified picture

Distribution

Polystichum world distribution map, present in Australia, China, and New Zealand

Identifiers

LSID urn:lsid:ipni.org:names:327121-2
WFO ID wfo-4000030832
COL ID 63P6K
BDTFX ID 87043
INPN ID 196554
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Hemigonum Cryptostigma Polystichum Hemesteum Porpaea Cyrtogonellum Cyrtomidictyum Acropelta Aetopteron Papuapteris Sorolepidium Hypopeltis Plecosorus

Lower taxons

Polystichum x kasayamense Polystichum aculeolatum Polystichum annapurnicola Polystichum bolanicum Polystichum calderonense Polystichum caudatum Polystichum caudescens Polystichum chaparense Polystichum cheilanthoides Polystichum copelandii Polystichum crinulosum Polystichum diaphanum Polystichum x eberlei Polystichum excelcior Polystichum x flemingii Polystichum x hagenahii Polystichum x hokurikuense Polystichum x iidanum Polystichum inadae Polystichum x inayatii Polystichum x jamunae Polystichum x jitaroi Polystichum x kai-montanum Polystichum x luerssenii Polystichum macleaii Polystichum maderense Polystichum magnificum Polystichum marionense Polystichum martini Polystichum x meyeri Polystichum papuanum Polystichum polyodon Polystichum x safarovii Polystichum x saltum Polystichum x tokyoense Polystichum eriorachis Polystichum x pseudosemifertile Polystichum bradei Polystichum brassii Polystichum cystostegium Polystichum dangii Polystichum kilimanjaricum Polystichum x kumamontanum Polystichum x kuratae Polystichum x microlepis Polystichum x midoriense Polystichum x minamitanii Polystichum x miuranum Polystichum moorei Polystichum x namegatae Polystichum oculatum Polystichum pseudolanceolatum Polystichum x pseudolentum Polystichum pseudomicrophyllum Polystichum x pseudoparvipinnulum Polystichum revolutum Polystichum x shin-tashiroi Polystichum x stewartii Polystichum x suginoi Polystichum x suyamanum Polystichum x takaosanense Polystichum x tare-bhirense Polystichum x tetsuyamense Polystichum x titibuense Polystichum x utsumii Polystichum alpinum Polystichum holttumii Polystichum cuneatum Polystichum keysserianum Polystichum kinabaluense Polystichum kurokawae Polystichum lancilobum Polystichum laniceps Polystichum kenwoodii Polystichum flaccidum Polystichum fuscum Polystichum volkensii Polystichum vulcanicum Polystichum walkerae Polystichum shizuokaense Polystichum spongiosum Polystichum stokesii Polystichum subtripteron Polystichum takakii Polystichum tortuosum Polystichum wawranum Polystichum weimingii Polystichum uahukaense Polystichum moluccense Polystichum mashikoi Polystichum muticum Polystichum rochaleanum Polystichum scariosum Polystichum rapense Polystichum meeboldii Polystichum peishanii Polystichum jiucaipingense Polystichum minutissimum Polystichum faucicola Polystichum speluncicola Polystichum centronepalense Polystichum palniense Polystichum x pseudobraunii Polystichum cavernicola Polystichum andersonii Polystichum dudleyi Polystichum haleakalense Polystichum kruckebergii Polystichum pungens Polystichum dracomontanum Polystichum macleae Polystichum setigerum Polystichum distans Polystichum erythrosorum Polystichum fournieri Polystichum furfuraceum Polystichum hartwegii Polystichum mickelii Polystichum muenchii Polystichum muricatum Polystichum platyphyllum Polystichum x potteri Polystichum kwakiutlii Polystichum speciosissimum Polystichum cochleatum Polystichum acutidens Polystichum erosum Polystichum woronowii Polystichum kadyrovii Polystichum retrosopaleaceum Polystichum mehrae Polystichum piceopaleaceum Polystichum omeiense Polystichum drepanum Polystichum falcinellum Polystichum dubium Polystichum craspedosorum Polystichum formosanum Polystichum hancockii Polystichum lepidocaulon Polystichum longifrons Polystichum obai Polystichum otomasui Polystichum polyblepharum Polystichum mayebarai Polystichum yaeyamense Polystichum montevidense Polystichum bakerianum Polystichum prescottianum Polystichum harpophyllum Polystichum subinerme Polystichum zambesiacum Polystichum mexiae Polystichum lehmannii Polystichum kungianum Polystichum woodsioides Polystichum machaerophyllum Polystichum excellens Polystichum fimbriatum Polystichum ichangense Polystichum kwangtungense Polystichum lanceolatum Polystichum leveillei Polystichum martinii Polystichum biaristatum Polystichum longipaleatum Polystichum prionolepis Polystichum hecatopterum Polystichum punctiferum Polystichum plaschnickianum Polystichum wilesianum Polystichum harrisii Polystichum underwoodii Polystichum ambiguum Polystichum trapezoides Polystichum mucronatum Polystichum christianae Polystichum echinatum Polystichum lemmonii Polystichum microchlamys Polystichum scopulinum Polystichum maevaranense Polystichum kalambatitrense Polystichum tsaratananense Polystichum coursii Polystichum rhizophyllum Polystichum triangulum Polystichum mohrioides Polystichum californicum Polystichum fragrans Polystichum andinum Polystichum fuentesii Polystichum tetragonum Polystichum bulbiferum Polystichum x bichnellii Polystichum tripteron Polystichum lentum Polystichum xiphophyllum Polystichum nepalense Polystichum manmeiense Polystichum neolobatum Polystichum semifertile Polystichum squarrosum Polystichum yunnanense Polystichum balansae Polystichum ovatopaleaceum Polystichum pseudomakinoi Polystichum rigens Polystichum tagawanum Polystichum thomsonii Polystichum bonapartii Polystichum stuebelii Polystichum acanthophyllum Polystichum alcicorne Polystichum atkinsonii Polystichum auriculum Polystichum basipinnatum Polystichum bifidum Polystichum chingae Polystichum christii Polystichum consimile Polystichum duthiei Polystichum fraxinellum Polystichum glaciale Polystichum nanchurnicum Polystichum setillosum Polystichum shensiense Polystichum sinense Polystichum subdeltodon Polystichum submite Polystichum yuanum Polystichum integripinnum Polystichum parvipinnulum Polystichum ningshenense Polystichum cyclolobum Polystichum wattii Polystichum decoratum Polystichum rhizophorum Polystichum tridens Polystichum turrialbae Polystichum paramoupinense Polystichum liui Polystichum jinfoshanense Polystichum parvifoliolatum Polystichum rupicola Polystichum deminuens Polystichum crinigerum Polystichum stimulans Polystichum shandongense Polystichum subacutidens Polystichum daguanense Polystichum paradeltodon Polystichum dielsii Polystichum tangmaiense Polystichum subulatum Polystichum gymnocarpium Polystichum x gemmilachenense Polystichum x silviamontanum Polystichum plicatum Polystichum aculeatum Polystichum pycnolepis Polystichum subfimbriatum Polystichum incongruum Polystichum pycnopterum Polystichum disjunctum Polystichum articulatipilosum Polystichum crassinervium Polystichum guangxiense Polystichum oblongum Polystichum assurgentipinnum Polystichum houchangense Polystichum muscicola Polystichum elevatovenosum Polystichum falcatilobum Polystichum jiulaodongense Polystichum zayuense Polystichum costularisorum Polystichum vestitum Polystichum pilosum Polystichum elmeri Polystichum schizophyllum Polystichum marquesense Polystichum albomarginatum Polystichum amboroense Polystichum bachii Polystichum chaparensis Polystichum congestum Polystichum lepidotum Polystichum paramicola Polystichum platylepis Polystichum giganteum Polystichum tijucense Polystichum manickamianum Polystichum australiense Polystichum fallax Polystichum formosum Polystichum whiteleggei Polystichum bonseyi Polystichum incisopinnulum Polystichum diffundens Polystichum exauriforme Polystichum deflexum Polystichum fugongense Polystichum fuscopaleaceum Polystichum acutipinnulum Polystichum anomalum Polystichum longissimum Polystichum sinotsussimense Polystichum pseudosetosum Polystichum levingei Polystichum oreodoxa Polystichum adungense Polystichum discretum Polystichum mucronifolium Polystichum carvifolium Polystichum nayongense Polystichum x lonchitiforme Polystichum rufopaleaceum Polystichum saxicola Polystichum altum Polystichum longispinosum Polystichum robustum Polystichum putuoense Polystichum fengshanense Polystichum australium Polystichum paleatum Polystichum submucronatum Polystichum proliferum Polystichum bigemmatum Polystichum otophorum Polystichum grandifrons Polystichum integrilobum Polystichum wulingshanense Polystichum cuneatiforme Polystichum submarginale Polystichum hagiangense Polystichum pseudotsus-simense Polystichum mulunense Polystichum rufum Polystichum chilense Polystichum subintegerrimum Polystichum hillebrandii Polystichum ammifolium Polystichum deltodon Polystichum hookerianum Polystichum quangbinhense Polystichum duyunense Polystichum tenuius Polystichum trapezoideum Polystichum minimum Polystichum simile Polystichum x amboversum Polystichum x hitoyoshiense Polystichum conjunctum Polystichum x rupestris Polystichum xichouense Polystichum uniseriale Polystichum guajaibonense Polystichum sanchezii Polystichum hainanicola Polystichum perpusillum Polystichum tonkinense Polystichum tiankengicola Polystichum normale Polystichum x kunioi Polystichum bicknellii Polystichum illyricum Polystichum x sarukurense Polystichum concinnum Polystichum talamancanum Polystichum huashanicola Polystichum mengziense Polystichum langchungense Polystichum taizhongense Polystichum pseudoacutidens Polystichum solomonii Polystichum bamlerianum Polystichum multifidum Polystichum obliquum Polystichum ordinatum Polystichum lachenense Polystichum rachichlaena Polystichum nudicaule Polystichum x illiricum Polystichum luctuosum Polystichum makinoi Polystichum alfaroi Polystichum x anceps Polystichum aquifolium Polystichum archboldii Polystichum x fominii Polystichum x fujisanense Polystichum x lesliei Polystichum lindseaefolium Polystichum lineare Polystichum longecuspis Polystichum neozelandicum Polystichum obtusum Polystichum x ohtanii Polystichum x okanum Polystichum x ongataense Polystichum wirtgenii Polystichum x yokohamense Polystichum silvaticum Polystichum shimurae Polystichum excelsius Polystichum bissectum Polystichum maximum Polystichum puteicola Polystichum guadalupense Polystichum monticola Polystichum transkeiense Polystichum transvaalense Polystichum igaense Polystichum ohmurae Polystichum imbricans Polystichum braunii Polystichum orbiculatum Polystichum lonchitis Polystichum munitum Polystichum setiferum Polystichum acrostichoides