Plants large, frequently covering extensive areas, dark green; saxicolous or corticolous, rarely terricolous. Primary stems creeping; secondary stems erect, to 120 mm long, stipe to 60 mm long, branches dense above stipe, branching simple to pinnate; in section round, central strand small and dense in erect stems, inner cortical cells small, thin-walled, yellowish, outer cortical cells smaller, ± thick-walled, brown. Leaves flattened, spreading, weakly crisped when dry; stem leaves broadly ovate to oblong-ovate, (1.0-)1.5-2.5(-3.0) mm long; acute to broadly acute; narrowed to insertion; margins plane, dentate, teeth multicellular; branch leaves similar to stem leaves but somewhat smaller. Costa single, extending to midleaf or above, smooth but terminating in a single dorsal tooth; in section bulging dorsally, in 3 rows of thickened, undifferentiated cells. Upper laminal cells irregularly rhomboidal, somewhat sigmoid, homogeneous, 35-50 µm long, 8-12 µm wide, walls weakly thickened, smooth; basal cells rectangular to rhomboidal, 40-75 µm long, 8-12 µm wide, walls weakly thickened, smooth; alar cells not differentiated. Perigonia axillary on branches, gemmate; perigonial leaves ovate, cuspidate, 1.0 mm long. Perichaetia along branches, obvious, green; perichaetial leaves oblong-acuminate, sheathing below, spreading at tips, to 2.5 mm long, leaf cells rhomboidal, thickened. Seta 15-25 mm long, red-brown, smooth. Capsule long-exserted, erect to inclined, ovoid, 2-3 mm long, smooth, red-brown, urn cylindrical, neck not differentiated; exothecial cells quadrate to rectangular or hexagonal, walls thickened, cells at mouth quadrate, strongly thickened, darker. Peristome double, yellow-brown; exostome teeth triangular, inflexed wet, erect dry, striate with median zigzag line, to 500 µm high; endostome segments above high basal membranes lanceolate and perforated or broad and cleft with diverging tips, shorter than teeth, weakly granulate; cilia present, linear, shorter than segments, weakly granulate. Operculum rostrate, 1 mm long. Calyptra cucullate, 2 mm long, smooth. Spores rounded, 9-15 µm, weakly granulate, brownish.
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