Dioecious or rarely monoecious, tendriled vines or lianas; stems 5-to 10-sulcate. Leaves petiolate, entire, lobate, or 3-to 5-foliolate; tendrils simple. Staminate flowers clustered at the end of long peduncles; calyx elongate cylin-dric, 5-lobate; corolla rotate, 5-lobate; stamens 2, included, free, the filaments short, the anthers linear or oblong, dorsifixed, bilocular, the theca straight, rarely replicate at the base. Pistillate flowers 1-3, short pedunculate; calyx and corolla as in the staminate flowers; ovary ovoid, 2-carpellate, the ovules numerous, hori-zontal, the styles connate on most of their length, the stigmas 2-branched; sta-minodia 2, minute. Fruits indehiscent, ovoid to oblong; seeds numerous, oblong, compressed.