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Marginal flowers often larger and with more unequal corolla lobes, inner flowers often smaller and more regular.
Corolla usually 5–fid or with 4–6–fid flowers in the same head, rarely entirely 4–fid.
Capitula hemispherical or globular, with 1–2 rows of foiaceous receptacular bracts.
Involucel ± sulcate, 8 furrowed, toothed or with a ± membranous corona.
Calyx short with 5–24 plumose setae of equal length.
Stigma oblique, entire or slightly 2 lobed.
Annual or perennial herbs or subshrubs.
Receptacle scales hairy or 0.
